Cryptic species of freshwater sculpin (Cottidae: Cottus) in California

The Riffle Sculpin (Cottus gulosus) is a small, bottom-dwelling fish regarded as widespread in the cool-water streams that flow into California’s Central Valley and into streams of the central California coast. Using population genomics, supported by other genetic, distributional, and meristic studies, we demonstrate that C. gulosus consists of three species with four subspecies (five lineages), all but one entirely endemic to California:
 
Cottus pitensis, Pit Sculpin Bailey and Bond 1963
Cottus gulosus, Inland Riffle Sculpin (Girard 1854)
            C. g. gulosus: San Joaquin Riffle Sculpin (Girard 1854), nominate subspecies
            C. g. wintu: Sacramento Riffle Sculpin, Moyle, Baumsteiger and Campbell 2022, new subspecies
Cottus ohlone, Coastal Riffle Sculpin Moyle, Baumsteiger and Campbell 2022, new species
            C. o. ohlone, Ohlone Riffle Sculpin Moyle, Baumsteiger and Campbell 2022, nominate subspecies
            C. o. pomo, Pomo Riffle Sculpin Moyle Baumsteiger and ...

溪杜父鱼（Cottus gulosus）是一种小型底栖鱼类，广泛分布于汇入加利福尼亚中央谷地的冷水溪流及加利福尼亚中部海岸的溪流中。借助群体基因组学（population genomics）技术，并辅以遗传、分布及可数性状研究（meristic studies），我们证实C. gulosus包含3个物种及4个亚种（共5个谱系），除其中一个外，其余均为加利福尼亚特有种：

Cottus pitensis，皮特杜父鱼（Bailey and Bond 1963）
Cottus gulosus，内陆溪杜父鱼（Girard 1854）
C. g. gulosus：圣华金溪杜父鱼（Girard 1854），指名亚种
C. g. wintu：萨克拉门托溪杜父鱼，Moyle、Baumsteiger及Campbell 2022年命名的新亚种
Cottus ohlone，海岸溪杜父鱼，Moyle、Baumsteiger及Campbell 2022年命名的新物种
C. o. ohlone：奥隆尼溪杜父鱼，Moyle、Baumsteiger及...