Intermediate habitat fragmentation buffers droughts: How individual energy dynamics mediate mammal community response to stressors

Biodiversity is threatened by land-use and climate change. Although these processes are known to influence species survival and diversity, predicting their combined effects on communities remains challenging. We here aim to disentangle the combined effects of drought-induced resource shortage and habitat fragmentation on species coexistence. To understand how both fragmentation and droughts affect individual movement and physiology, and ultimately influence population and community dynamics, we use an individual-based metabolic modelling approach to simulate a community of small mammals. Individuals forage in the landscape to ingest energy, which they then allocate to basal maintenance, digestion, locomotion, growth, reproduction, and storage. If individuals of several species are able to balance their energy intake and needs, and additionally store energy as fat reserve, they may overcome stress periods and coexist. We find that species recover best after a drought when they live in mo..., In this study, we used a relatively novel dynamic individual-based metabolic simulation model for a mammal community (see also https://doi.org/10.5061/dryad.4qrfj6qjf). The model is based on allometric relationships and movement in home ranges, allowing for a variety of species. The model was thoroughly tested and validated using real-world patterns from the literature. An extensive model development description is available in the format of a TRACE document with the publication. We used the model to simulate scenarios of different habitat fragmentation in combination with drought events and analyzed the resulting data., # Intermediate habitat fragmentation buffers droughts: How individual energy dynamics mediate mammal community response to stressors

We here provide the model code for our individual-based metabolic community model along with the resulting data for the simulated experiments and the code for producing the manuscript figures for \"Intermediate habitat fragmentation buffers droughts: How individual energy dynamics mediate mammal community response to stressors\"

### Files

The file \"UFZ_Drought_Monitor_daily_SM_L02_subset.nc\" contains data of the UFZ Drought Monitor / Helmholtz Centre for Environmental Research. It is a subset of the daily Soil Moisture Index (SMI) data from [https://www.ufz.de/index.php?en=37937](https://www.ufz.de/index.php?en=37937) for two time periods and 20 random locations, which were used to simulate observed droughts with the model. The data was provided by Dr. Andreas Marx and is based on Zink et al. 2015 (DOI 10.1088/1748-9326/11/7/074002).

The achive \"mibcom_d...,

生物多样性正遭受土地利用变化与气候变化的双重威胁。尽管已知这两类过程均会影响物种存活与群落多样性，但精准预测二者对生物群落的联合效应仍颇具挑战。本研究旨在厘清干旱引发的资源短缺与生境破碎化对物种共存的联合作用机制。为明晰生境破碎化与干旱如何分别影响个体运动与生理机能，并最终作用于种群及群落动态，我们采用基于个体的代谢建模（individual-based metabolic modelling）方法，模拟小型哺乳动物群落。个体在景观中觅食以获取能量，并将能量分配至基础维持、消化、运动、生长、繁殖与储能环节。若多个物种种群能够实现能量摄入与需求的平衡，且可将过剩能量以脂肪储备形式储存，则可顺利度过胁迫期并实现稳定共存。我们发现，当物种栖息于适度……（原文此处截断为mo...）。

本研究针对哺乳动物群落采用了相对新颖的动态基于个体的代谢模拟模型（dynamic individual-based metabolic simulation model）（详见https://doi.org/10.5061/dryad.4qrfj6qjf）。该模型基于异速生长关系与家域运动规律，可适配多样物种种群。模型已通过已发表文献中的真实观测数据完成全面测试与验证，详尽的模型开发说明以配套TRACE文档形式随正式发表论文同步公开。我们借助该模型模拟了不同生境破碎化程度配合干旱事件的多组场景，并对模拟产生的数据集开展系统性分析。
# 中等程度生境破碎化可缓解干旱胁迫：个体能量动态如何介导哺乳动物群落对胁迫因子的响应
本研究提供基于个体的代谢群落模型（individual-based metabolic community model）的源代码，以及模拟实验产生的配套数据集，同时包含用于绘制论文插图的代码，对应论文标题为《中等程度生境破碎化可缓解干旱胁迫：个体能量动态如何介导哺乳动物群落对胁迫因子的响应》。

### 数据文件
文件"UFZ_Drought_Monitor_daily_SM_L02_subset.nc"包含德国亥姆霍兹环境研究中心（UFZ）干旱监测数据集。该数据集是每日土壤湿度指数（Soil Moisture Index, SMI）数据的子集，源自[https://www.ufz.de/index.php?en=37937]，涵盖两个时间段与20个随机采样点位，用于驱动模型模拟观测到的干旱事件。该数据由Andreas Marx博士提供，其原始研究基于Zink等人2015年的工作（DOI: 10.1088/1748-9326/11/7/074002）。

归档文件"mibcom_d..."（原文此处截断）