Description of the thirteen loci used in this study averaged across all sites.

Ta = annealing temperature, Ho = average observed heterozygosity, He = expected average heterozygosity, P = significance value from comparison between Ho and He, r = frequency of null alleles, As = average number of alleles per collection site, A = total alleles observed at each locus.1Non-species-specific markers.†weakly significant (0.05≥P≥0.01).