A complete mandible of a new amphisbaenian reptile (Squamata, Amphisbaenia) from the late middle eocene (Bartonian, Mp 16) of France

ABSTRACTA new genus of amphisbaenian reptile, <i>Cuvieribaena carlgansi</i>, is decribed from the late middle Eocene of France. This material from the locality of Le Bretou (Phosphorites du Quercy) is an almost completely preserved mandible. The dentary bears six teeth, with the anterior-most one the largest. The anterior teeth are blunt and enlarged (‘amblyodont’), and their size and amblyodonty decrease posteriorly. The presence of well-developed amblyodont teeth indicates a durophagous diet. Durophagy has already been recognized in some amphisbaenids, and their high skull, short jaws, and robust tooth morphology predispose them to this durophagous lifestyle. Several other amphisbaenid specimens have also been recovered from Le Bretou, mainly trunk vertebrae and two incomplete dentaries. This material and the complete mandible show unambiguous morphological features tying them to Blanidae. <i>Cuvieribaena</i> is the earliest secure record of blanid lizards in the fossil record, confirmed by phylogenetic analysis.http://zoobank.org/urn:lsid:zoobank.org:pub:17454922-A121-447F-9D01-E4992ADB7D60SUPPLEMENTAL DATA—Supplemental materials are available for this article for free at www.tandfonline.com/UJVP

摘要：本文记述了产自法国始新世中晚期的蚓蜥类爬行动物（amphisbaenian reptile）新属新种*Cuvieribaena carlgansi*。该标本采自勒布勒图（Le Bretou）产地（凯尔西磷灰岩，Phosphorites du Quercy），为一件近乎完整的下颌骨。齿骨上保留有6枚牙齿，最前端的牙齿体型最大。前部牙齿钝圆且膨大（amblyodont，钝齿型），其尺寸与钝齿特征均向后逐渐减弱。发达的钝齿型牙齿表明该物种具备食硬食物性（durophagy）的食性。食硬食物性在部分蚓蜥科物种中已有记录，这类类群高耸的颅骨、短缩的下颌以及粗壮的牙齿形态均使其适配此类食性。此外，研究团队还从勒布勒图地点出土了其他多件蚓蜥类标本，以躯干椎骨和两件不完整的齿骨为主。这批标本与完整下颌骨均具有明确的形态特征，可归入布兰蜥科（Blanidae）。*Cuvieribaena*是化石记录中布兰蜥科蜥蜴最确凿的最早化石记录，该结论得到了系统发育分析的证实。本文对应的Zoobank注册号为http://zoobank.org/urn:lsid:zoobank.org:pub:17454922-A121-447F-9D01-E4992ADB7D60。补充数据：本文补充材料可于www.tandfonline.com/UJVP免费获取。