Campichoeta (Campichoeta) spinicauda Papp, 2005, sp. n.

Campichoeta (Campichoeta) spinicauda sp. n. (Figs 23–25) Holotype male (HNHM): TAIWAN: Taipei Hsien, Fu-Shan LTER Site, lake shore, March 25, 2003, No. 3 – a meadow & dry bed of a creek, leg. M. FÖLDVÁRI. Paratypes: 1 female (HNHM): ibid., Kaohsiung Hsien, Liukuei, Shan-Ping LTER Site, over/along a creek, April 2, 2003, No. 18, leg. M. FÖLDVÁRI; 1 male (NMNS, glued on tip of a point label, damaged, head and wings broken down and glued to label below minuten, gen. prep. in a plastic microvial with glycerol): Taiwan Nantou, Hsini Tungpu 1/II/1989, K. W. HUANG, Sweeping net – 431–482 [NMNS id. label]. Description – Measurements in mm: body length 2.86 (holotype), 2.65, 3.30 (paratypes), wing length 2.62 (holotype), 2.45, 2.86 (paratypes), wing breadth 0.90, 0.88, 0.98. Dark graphite grey with thick brownish grey microtomentum (head, thorax, abdomen). Cephalic setae as in C. obscuripennis: a more laterally placed proclinate pair of frontoorbitals, a large reclinate ors just behind and medially to the proclinate one and a third short thin reclinate ors anteriorly and medially to the proclinate ors. Ocellars, inner and outer verticals very long, postocellars long (0.22 mm), cruciate. Frons with some small setae behind lunule. Vibrissae comparatively short but thick. Gena only 0.025 mm broad, 5–6 thin, 0.065–0.07 mm long peristomals present. Subgenal seta thin, 0.11 mm long. Antenna and arista black. First flagellomere 0.31 mm long (measured dorsally), 0.12 mm broad, covered by 0.02 mm long cilia, longest (dorsal) aristal cilia 0.025 mm. Mesonotum moderately humped. Thoracic setae: 1 pprn, 2 np, 1 large prst, 0+2 dc, 1 large prealar, 1 sa, 1 smaller pa, 1 prsc, 2 sc; 1 dorsal and 1 ventral large kepst. mm for Figs 25–26, 0.1 mm for Figs 21–24 Wing uniformly dark brown, veins even darker (almost black). Longest costal setae distally to H (mg1) 0.07 mm, 0.055 mm on mg2, the longer seta at break 0.11 mm. M vein sections 0.715, 0.97, ratio 1.35. Terminal section of Cu 0.33 mm. Anal cell very small. Anal (A1) and axial (A2) veins present as shadows of veins only. Squamal cilia black. Halteres greyish yellow. Legs mainly yellow; coxae and femora (except yellow apical 1/5) greyish brown. Fore femur posteroventrally with 7 (paratype: 8) peg-like black thornlets. Fore coxa with 2 long ventrally curved and some other shorter setae. Mid femur with a row of long anterior setae. Preapicals on tibiae comparatively weak, ventroapical of mid tibia 0.16 mm long. Sternites (Fig. 25) similar to those of C. obscuripennis (Fig. 26), i.e. long and narrow but with distinctly fewer setae: sternite 4 with 12 (vs 20), sternite 5 with 14 (vs 22) setae. The holotype male with 4, the paratype male with 6 long setae on tergite 7 (medial pair of the paratype must be weaker, judged by the setal bases). Male surstylus reduced, or rather, fused to epandrium as its ventromedial lobe (Figs 21, 23). This is one of the main distinctive features of the nominate subgenus (males of the subgenus Thryptocheta Rondani have true surstyli). Surstylar lobe (Fig. 23) more slender, distinctly less broad than in C. obscuripennis (Fig. 21), and with less setae. Gonopod (Fig. 24) long and slender as in obscuripennis (Fig. 22) but definitely different (when seen in its broadest extension). Female cerci short, hidden under tergite 7 and tergite 8, with short thin setae only. Remarks – This new species does not only belong to the C. obscuripennis group (i.e. to the nominate subgenus Campichoeta), also its closest relative is C. obscuripennis (MEIGEN, 1830). The new species is different from C. obscuripennis by the number of setae on abdominal sternites and by the details of the male genitalia (see above and in figures). OKADA (1966) reported C. obscuripennis from Nepal. Now I question the identity of that Nepalese species.棘尾坎姬蝇（Campichoeta (Campichoeta) spinicauda）新种（图23~25） 正模雄性（匈牙利自然历史博物馆（HNHM））：采自中国台湾省台北县福山长期生态研究站湖岸，2003年3月25日，编号3，生境为草地与干涸溪床，采集人M. FÖLDVÁRI。 副模：1头雌性（匈牙利自然历史博物馆（HNHM））：采自中国台湾省高雄县六龟乡山排长期生态研究站溪上及溪岸，2003年4月2日，编号18，采集人M. FÖLDVÁRI；1头雄性（国立自然科学博物馆（NMNS），标本粘于针标末端，头部与翅膀破损并粘于微针下方的标签上，生殖器制备于含甘油的塑料微型小管中）：采自中国台湾省南投县仁里东埔，1989年2月1日，采集人K. W. HUANG，采集方式为网扫，编号431~482，附NMNS鉴定标签。 形态测量（单位：mm）：体身长2.86（正模）、2.65、3.30（副模）；翅长2.62（正模）、2.45、2.86（副模）；翅宽0.90、0.88、0.98。全体呈深石墨灰色，被厚密的棕灰色微粉被（覆盖头部、胸部与腹部）。 头部鬃毛特征与暗翅坎姬蝇（C. obscuripennis）一致：1对侧位的前倾额眶鬃，1根粗大的后倾眶鬃位于前倾额眶鬃的后内侧，另有3根短小纤细的后倾眶鬃位于前倾额眶鬃的前内侧。单眼鬃、内顶鬃与外顶鬃极长，后顶鬃长0.22 mm，交叉排列。额部在新月片后方具若干小鬃。触须相对较短但粗壮。颊部宽仅0.025 mm，具5~6根细长的缘鬃，长0.065~0.07 mm。亚颊鬃纤细，长0.11 mm。触角与触角芒呈黑色。第一鞭节长0.31 mm（背侧测量），宽0.12 mm，被长0.02 mm的纤毛覆盖，背侧最长的芒纤毛长0.025 mm。 中胸背板适度隆起。胸部鬃序：1根前背侧鬃、2根背侧鬃、1根粗大的前盾鬃、0+2根中鬃、1根粗大的前翅后鬃、1根翅上鬃、1根较小的后翅上鬃、1根后盾鬃、2根小盾鬃；1根背侧与1根腹侧的大型后侧片鬃。 图25~26比例尺为1 mm，图21~24比例尺为0.1 mm。 翅呈均匀深棕色，翅脉颜色更深（近黑色）。H脉（mg1）远端的最长缘脉鬃长0.07 mm，mg2处的缘脉鬃长0.055 mm，翅脉断点处的较长鬃长0.11 mm。M脉各段长分别为0.715、0.97，比例为1.35。Cu脉末端段长0.33 mm。臀室极小。臀脉（A1）与轴脉（A2）仅呈脉状阴影。翅瓣纤毛黑色。平衡棒呈灰黄色。 足主要呈黄色；基节与股节（除顶端1/5为黄色外）呈灰棕色。前足股节后腹侧具7根（副模为8根）黑色钉状小刺。前足基节具2根向腹侧弯曲的长鬃，以及若干短鬃。中足股节具1列长的前侧鬃。胫节的端前鬃相对较弱，中足胫节的腹端鬃长0.16 mm。 腹部腹板（图25）与暗翅坎姬蝇（图26）相似，即狭长但鬃毛数量明显更少：第4腹板具12根鬃（对比暗翅坎姬蝇的20根），第5腹板具14根鬃（对比暗翅坎姬蝇的22根）。正模雄性第7背板具4根长鬃，副模雄性具6根长鬃（根据鬃毛基部判断，副模的内侧对鬃相对较弱）。 雄性尾须退化，更确切地说，与下生殖板愈合为其腹内侧叶（图21、23）。这是指名亚属的主要鉴别特征之一（Thryptocheta亚属的雄性具真正的尾须）。尾须叶（图23）更为细长，明显比暗翅坎姬蝇（图21）的更窄，且鬃毛更少。生殖肢（图24）与暗翅坎姬蝇（图22）一样细长，但在最宽处观察时存在明显差异。 雌性尾须短，隐藏于第7、8背板下方，仅具短小纤细的鬃毛。 讨论：本新种隶属于暗翅坎姬蝇种团（即指名坎姬蝇亚属），其近缘种为暗翅坎姬蝇（MEIGEN, 1830）。本新种与暗翅坎姬蝇的区别在于腹部腹板的鬃毛数量以及雄性外生殖器的细节（详见前文及图版）。OKADA（1966）曾报道采自尼泊尔的暗翅坎姬蝇，现笔者对该尼泊尔标本的物种鉴定存疑。