Manota bilobata Papp, 2004, sp. n.

Manota bilobata sp. n. (Figs 13–17) Holotype: male, TAIWAN: Kaohsiung Hsien, Liukuei, Shan-Ping LTER Site – creek valley, No.13, Mar 31-Apr 1, 2003, L. Papp & M. Földvári. (HNHM, abdomen with genitalia in a plastic microvial with glycerol, otherwise intact.) Paratypes: 1 female (HNHM, left mid leg, right hind tarsomeres 2–5 lost): ibid., UV light traps, March 31– April 4, No. 14, M. Földvári & L. Papp; 1 female (HNHM, right hind leg, left fore tarsomeres 2–5, left hind tarsomeres 3–5 and right flagellum lost): TAIWAN: Taipei Hsien, Fu-Shan LTER Site, No. 7 – lake shore, UV light, March 25–28, 2003, M. Földvári & L. Papp; 1 male (HNHM, damaged, mid and hind legs, right fore tibial spur and left flagellomeres 8–14 lost): TAIWAN: Taipei, Nanshih Chiao, Han-Lo-Da, 450 m – rocky forest undergrowth, Sep 23, 2000, leg. L. Papp, No. 1; 1 male (NMNS, left hind leg lost, prepared from alcohol, so body and wings wrinkled; abdomen with genitalia in a plastic microvial with glycerol): TAIWAN: Nantou, Yuchig Lienhunchi, IV-V/9–10/1998, C.S. Lin & W. T. Yang, Malaise trap – NMNS ENT 3028–971. Measurements in mm: body length 2.64 (holotype), 2.28, 2.70 (paratype males), 2.47, 3.10 (paratype females), wing length 2.26, 2.00, 2.20 (paratype males), 2.28, 2.70 (paratype females), wing breadth ca. 0.85 (holotype), 0.85, not measurable, ca. 0.87 and 1.25 (paratypes). Thorax mostly ochre, caudal part of thorax incl. scutellum and mediotergite diffuse brown (one of the female paratypes with almost completely dark thorax). Eyes with evenly distributed medium-long hairs. Facial setae yellowish at middle, dark over clypeus and along eyes, or all facial setae dark. Postocellar setae black. Longest postocular setulae 0.065–0.09 mm. Scape, pedicel and flagellomeres dark. Flagellum of holotype ca. 1.25 mm long (much curved). Flagellomeral hairs dense, 0.025 mm, apically 0.035 mm long. Palpi white or whitish yellow, ca. 0.55 mm long on holotype, ca. 0.48 mm on one male, 0.75 mm long on one female paratype. Anepisternum with dense hairs, katepisternum pilose on anteroventral part (badly visible), laterotergite bare. Scutellum with 2 pairs of long black marginal setae. Wing light brownish with brown veins. Costal section from H to R 1 0.715 mm, from R 1 to R 5 1.01 mm, extant sections of M 1 0.485 mm (to wing margin), M 2 1.31 mm long. Halteres ochreous with blackish knob. Legs yellow. Length of femur, tibia, first tarsomere and tibial spurs (in mm, measured on holotype): fore: 0.605, 0.46, 0.50, 0.34, mid: 0.95, 0.95, 0.57, 0.42, 0.32; hind: 1.05, 1.36, 0.605, 0.46, 0.39. Abdomen dark brown, sternites 1–5 lighter. Postabdomen incl. segment 9 ochre. Tergite 9 longer than broad (Fig. 13), apically with some longer thick setae. Sternite 9 (Fig. 14) subtriangular, its setae are shorter but more numerous than on tergite 9. 0.2 mm for all Cerci, which bear some minute black thornlets apically. Gonocoxite (Figs 15–16) with a bulbous ventral lobe and with 2 long thick, apically curved black setae at the base of gonostylus (Fig. 17). Gonocoxites close to each other. Gonostylus long, setose, bilobed (Figs 15–17). Hypoproct with a pair of long but not thick setae and with several short setulae. Aedeagus short and triangular. Female body and wing length probably slightly larger than in males (see above). Postabdomen ochre, cerci with evenly distributed hair-like setae not longer than 0.035 mm. The females were identified by the body colour and by the colour of facial and fore coxal setae. M. bilobata sp. n. is easily separable from the other Taiwanese species of Manota by its black postocular setae and partly dark fore coxal setae. It belongs to that species-group of the true Manota, where the laterotergite is bare and the katepisternum is pilose (anepisternum is with dense hairs); in these respects it is similar to M. chinensis and M. delyorum. The distinctive features are in the male genitalia: The gonocoxite has only 2, not particularly thick megasetae near gonostylar base (cf. SØLI 1993), and the gonostylus is bilobed with both lobes bearing longer setae.双叶曼诺塔蚊（Manota bilobata sp. n.）（图13~17） 正模标本（Holotype）：雄虫，采自中国台湾高雄县六龟乡山坪长期生态研究站溪谷，编号13，2003年3月31日至4月1日，由L. Papp与M. Földvári采集。标本保存于匈牙利自然历史博物馆（HNHM），腹部及生殖器置于含甘油的塑料微型小管内，躯体其余部分完整。 副模标本（Paratypes）：1头雌虫副模（HNHM）：左侧中足缺失，右侧后足跗节第2~5节缺失，采集信息同正模，采用紫外灯诱捕（UV light traps），2003年3月31日至4月4日，编号14，由M. Földvári与L. Papp采集；1头雌虫副模（HNHM）：右侧后足缺失，左侧前足跗节第2~5节、左侧后足跗节第3~5节及右侧鞭节缺失，采自中国台湾台北县福山长期生态研究站湖岸，紫外灯诱捕，2003年3月25日至28日，编号7，由M. Földvári与L. Papp采集；1头雄虫副模（HNHM）：躯体受损，中足、后足、右侧前足胫距及左侧鞭小节第8~14节缺失，采自中国台湾台北县南势角汉罗大海拔450米处多岩森林下层灌丛，2000年9月23日，由L. Papp采集，编号1；1头雄虫副模（台湾自然科学博物馆，NMNS）：左侧后足缺失，由酒精保存液制备，躯体及翅褶皱；腹部及生殖器置于含甘油的塑料微型小管内，采自中国台湾南投县鱼池乡莲华池，1998年IV-V期/9~10日，由C.S. Lin与W.T. Yang采集，采用马氏陷阱（Malaise trap）采集，标本编号NMNS ENT 3028–971。 测量单位为毫米：体长：正模2.64，雄虫副模2.28、2.70，雌虫副模2.47、3.10；翅长：雄虫副模2.26、2.00、2.20，雌虫副模2.28、2.70；翅宽约0.85（正模），副模分别为0.85、无法测量、约0.87及1.25。 胸部整体呈赭黄色，胸部后侧区域（包括小盾片（scutellum）及中背板（mediotergite））具弥散状棕褐色斑纹，其中1头雌虫副模的胸部几乎完全呈黑色。复眼覆有均匀分布的中等长度毛。颜面鬃在中部呈淡黄色，在唇基（clypeus）及沿复眼处呈黑色，或全部颜面鬃均为黑色。单眼后鬃（postocellar setae）呈黑色。最长眼后鬃（postocular setulae）长0.065~0.09毫米。柄节（scape）、梗节（pedicel）及鞭小节（flagellomeres）均呈黑色。正模的鞭节长约1.25毫米，明显弯曲。鞭小节覆有密集的毛，毛长0.025毫米，端部毛长0.035毫米。触须（palpi）呈白色或淡黄白色，正模触须长约0.55毫米，1头雄虫副模的触须长约0.48毫米，1头雌虫副模的触须长0.75毫米。 前侧片（anepisternum）覆有密集的毛，后侧片（katepisternum）的前腹区域被毛（观察效果不佳），侧背片（laterotergite）无毛。小盾片具2对长而黑色的缘鬃。 翅呈淡棕褐色，翅脉为棕褐色。翅脉肋区：从肩横脉（H）到径脉1（R₁）长0.715毫米，从径脉1（R₁）到径脉5（R₅）长1.01毫米；中脉现存片段：M₁长0.485毫米（至翅缘），M₂长1.31毫米。平衡棒呈赭黄色，端球呈黑褐色。 足呈黄色。根据正模测量的各足股节、胫节、第1跗节及胫距（tibial spurs）长度（单位：毫米）：前足：0.605、0.46、0.50、0.34；中足：0.95、0.95、0.57、0.42、0.32；后足：1.05、1.36、0.605、0.46、0.39。 腹部呈深棕褐色，第1~5腹板（sternites）颜色较浅。后腹部（包括第9背板（tergite））呈赭黄色。第9背板长大于宽（图13），端部具若干较长的粗鬃。第9腹板（sternite 9，图14）呈亚三角形，其上的鬃较第9背板短但数量更多。尾须（cerci）长均为0.2毫米，端部具若干细小的黑色棘突。生殖基节（gonocoxite，图15~16）具球状腹叶，在生殖刺突（gonostylus，图17）基部具2根长而粗、端部弯曲的黑色鬃。生殖基节彼此靠近。生殖刺突细长，被毛，分为两叶（图15~17）。下生殖板（hypoproct）具1对长但不粗壮的鬃，以及若干短毛。阳茎（aedeagus）短而呈三角形。 雌虫的体躯及翅长略大于雄虫（详见前文）。后腹部呈赭黄色，尾须覆有均匀分布的毛状鬃，毛长不超过0.035毫米。雌虫根据体躯颜色、颜面及前足基节鬃的颜色进行鉴定。 双叶曼诺塔蚊（M. bilobata sp. n.）可通过其黑色的眼后鬃以及部分呈黑色的前足基节鬃，与台湾地区其他曼诺塔蚊属（Manota）物种轻易区分。本种隶属于真曼诺塔蚊类群，该类群的侧背片无毛、后侧片被毛（前侧片覆有密集的毛）；在这些特征上，本种与中华曼诺塔蚊（M. chinensis）及德氏曼诺塔蚊（M. delyorum）相似。其鉴别特征主要在于雄虫外生殖器：生殖基节仅在生殖刺突基部具2根（并非特别粗壮的）大鬃（megasetae，参照SØLI 1993），且生殖刺突分为两叶，两叶均具较长的鬃。