Frequency of observed behavioural measures for each experimental treatment.

These include: a) the frequency of cleaning interactions, b) mean inspection duration, c) proportion of interactions in which tactile stimulation was applied to clients, d) frequency of jolts per 100 s of inspection, e) frequency of chases, f) duration of chases and g) frequency of bites given by focal individual. Mean ± Standard Error (SEM) are provided for each behavioural measure.