Contributions of variables in models relating three metrics of invertebrate assemblages (assemblage structure, total catch of individuals, species richness) to experimental trampling, key habitat attributes (wrack cover, sediment compactness, grain size) and the time pitfall traps were deployed during the experiments.

Variable contributions are assessed in two complementary ways: i) a multi-model inference approach using cumulative weights, w+(j), and ii) the proportion of variance explained in distance-based linear models.