Table 4 in Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java

Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold.
D (in mm)H (in mm)whorlsD/HU/Dshell peripheryshell colorationscale sculpturespiral lines on apical sidepenispenis coecumflagellumnode at flagellumL. winteriana9.7–12.6 (mean 11.8)5.5–7.8 (mean 6.7)5–5.75 (mean 5.3)1.55–2.07 (mean 1.76)0.29–0.51 (mean 0.35)keeledbrownish- corneousscaly processes sparse, at peripheryindistinctwith large globular bulge at proximal endabsentdistal part crenated, proximal part shortabsentL. conoidea7.6–8.7 (mean 8.1)5.2–5.6 (mean 5.3)4.75–5 (mean 4.9)1.48–1.54 (mean 1.52)0.30–0.33 (mean 0.31)sharply keeledbrownish- corneoustubercles all over teleoconchnot discernible????L. rotatoria15.6–19.1 (mean 17.4)8.3–10.6 (mean 9.5)5.75–6.5 (mean 6.1)1.57–2.09 (mean 1.85)0.24–0.32 (mean 0.29)keeledbrownscaly processes sparse, at peripheryindistinctwith small globular bulge at proximal endabsentlongish lemon-shapedabsentL. naggsi sp. nov.11.6–14.0 (mean 12.5)8.5–10.6 (mean 9.6)6.25–7 (mean 6.6)1.21–1.37 (mean 1.31)0.28–0.33 (mean 0.31)angularbrownish- corneousscaly processes all over teleoconchnot discernible????L. epiplatia11.8–17.0 (mean 15.3)5.7–6.9 (mean 6.0)5.25–6.25 (mean 5.7)2.03–2.94 (mean 2.52)0.30–0.43 (mean 0.37)sharply keeledbrownish- corneousscaly processes sparse, at peripheryindistinctvery long, with small bulge at proximal endabsentvery long, tapering towards proximal endabsentL. parahyangensis sp. nov.14.7–16.9 (mean 15.7)8.4–9.3 (mean 8.7)6.5–7 (mean 6.8)1.69–1.94 (mean 1.80)0.39–0.45 (mean 0.42)sharply keeledbrownishcorneousscaly processes sparse, at peripherynot discernible????L. nusakambangensis sp. nov.11.9–15.0 (mean 13.5)6.6–9.1 (mean 7.9)5.25–6 (mean 5.7)1.65–1.83 (mean 1.72)0.32–0.40 (mean 0.36)keeledbrownishcorneousscaly processes sparse, at peripheryindistinctwith small globular bulge at proximal endabsentdistal part slightly crenated, proximal part longabsentL. petrukensis sp. nov.8.1–9.8 (mean 9.0)5.2–6.3 (mean 5.5)4.75–5.5 (mean 5.1)1.49–1.79 (mean 1.62)0.24–0.37 (mean 0.27)sharply keeledbrownish- corneousscaly processes all over teleoconchindistinctdivided by narrowing into long, cylindrical distal part and slightly dilated proximal partabsentslightly crenated, abruptly tapering at proximal endabsentL. tholiformis sp. nov.13.0–14.4 (mean 13.9)7.4–7.8 (mean 7.7)6–6.25 (mean 6.1)1.77–1.88 (mean 1.82)0.32–0.40 (mean 0.35)sharply keeledbrownish- corneousscaly processes all over teleoconchindistinctslightly broader proximallyshortcrenated, slightly longer than epiphallusabsentL. abdidalem sp. nov.13.1–16.4 (mean 14.0)6.3–7.8 (mean 6.9)5.75–6 (mean 5.9)1.93–2.14 (mean 2.03)0.33–0.48 (mean 0.38)sharply keeledbrownish- corneousscaly processes sparse, at peripheryindistinctlong and narrow, without proximal buldgeabsentcrenated, shorter than epiphallusabsentlong, broadest inL. madurensis sp. nov.11.4–12.0 (mean 11.6)5.5–6.5 (mean 5.9)5.5–5.75 (mean 5.7)1.75–2.11 (mean 1.98)0.38–0.40 (mean 0.39)sharply keeledbrownish- corneousscaly processes protruding into hairsno spiral lineslong, divided into longer, broader distal and shorter, narrower proximal partabsentmiddle part, gradually tapering towards proximal end, slightly crenatedabsentL. sewuensis sp. nov.11.4–16.3 (mean 13.7)6.3–8.7 (mean 7.2)5.25–6 (mean 5.7)1.61–2.25 (mean 1.90)0.30–0.40 (mean 0.36)sharply keeledbrownish- corneousscaly processes sparse, at peripheryindistinctwith globular swelling at proximal endabsentbroad, with two rows of tuberclesabsentL. sukoliloensis sp. nov.10.4–12.3 (mean 11.2)6.0–7.4 (mean 6.5)5–5.75 (mean 5.5)1.60–1.93 (mean 1.72)0.22–0.41 (mean 0.29)sharply keeledbrownish- corneousscaly processes sparse, at peripheryindistinctslightly dilated at proximal endabsentabruptly tapering at proximal end, not distinctly crenatedabsentL. tonywhitteni sp. nov.14.3–17.6 (mean 16.2)7.0–8.5 (mean 7.5)5.75–6 (mean 5.8)1.93–2.34 (mean 2.15)0.32–0.41 (mean 0.37)sharply keeledbrownish- corneousradial scaly processesindistinctdivided into long, cylindrical distal part and dilated proximal partabsentshort, broadest at base, not distinctly crenatedpresentL. nodifera sp. nov.14.7–16.1 (mean 15.2)6.4–8.1 (mean 7.2)5–5.75 (mean 5.5)1.93–2.29 (mean 2.12)0.26–0.38 (mean 0.32)sharply keeledbrownish- corneousscaly processes sparse, at peripheryindistinctdivided into cylindrical distal and dilated proximal partpresentmoderately long, tuberculatedpresentL. pacitanensis sp. nov.10.6–14.7 (mean 13.1)6.2–8.2 (mean 7.1)5.25–6 (mean 5.7)1.57–2.18 (mean 1.84)0.28–0.34 (mean 0.31)sharply keeledbrownish- corneousscaly processes sparse, at peripheryindistinctdivided by narrowing into long distal and short proximal sectionpresentmoderately long, not crenatedabsentL. intumescens11.6–14.1 (mean 13.2)6.1–8.7 (mean 7.5)5–5.75 (mean 5.3)1.50–1.98 (mean 1.76)0.22–0.34 (mean 0.27)sharply keeledbrownish- corneouswithout scaly processesindistinctdivided by narrowing into long distal and short proximal sectionabsentdistal part crenated, proximal part shortabsentL. moussoniana10.2–12.0 (mean 11.1)5.5–6.7 (mean 6.3)5–6.25 (mean 5.7)1.73–1.81 (mean 1.77)0.37–0.38 (mean 0.38)keeledbrownish- corneous?indistinct????L. schepmani10.5–10.96.1–6.25–5.251.72–1.760.21sharply keeledbrownish- corneousno scales discernibledistinct????L. monticola8.5–9.5 (mean 9.0)5.3–6.0 (mean 5.8)4.5–5.25 (mean 4.8)1.49–1.60 (mean 1.56)0.18–0.25 (mean 0.21)keeledyellowish or brownish- corneousno scales discernibledistinct????slightly crenated,L. smimensis6.3–11.9 (mean 8.2)4.1–8.4 (mean 5.6)4.5–5.5 (mean 4.9)1.28–1.64 (mean 1.48)0.18–0.32 (mean 0.25)angular or roundedbrownish- corneoushair-like processesindistinctlong, gradually tapering towards its proximal endabsentgradually tapering towards proximal endabsentL. leucochila16.3–17.58.5–9.55.251.920.30angularyellowishscaly processes all over teleoconchindistinct????L. zonifera sp. nov.12.5–13.2 (mean 12.8)6.9–7.6 (mean 7.2)5.5–5.75 (mean 5.6)1.73–1.81 (mean 1.77)0.40keeledcorneous with brownish zone at peripheryscaly processes sparse, at peripheryindistinctslightly dilated at proximal endabsentslightly crenated, more than twice as long as epiphallusabsentL. ciliocincta12.4–15.6 (mean 13.6)6.0–8.3 (mean 7.0)5–6 (mean 5.4)1.86–2.12 (mean 1.94)0.22–0.30 (mean 0.27)keeledwhitish- corneous with brownish band at peripheryscaly processes sparse, at peripheryindistinctwith small bulge at proximal endabsentshort, very broad, tapering towards proximal endabsentmoderatelywhitish,long, almostL. pakidulan sp. nov.12.5–14.0 (mean 13.4)6.9–8.1 (mean 7.4)5.25–5.5 (mean 5.4)1.70–1.86 (mean 1.81)0.24–0.30 (mean 0.27)angularwith reddish- brown band atwithout scaly processesindistinctshort, with large bulge at proximal endabsentsmooth, gradually tapering nearabsentperipheryproximalendwhitish,withL. dharmai sp. nov.9.97.25.51.390.18angularreddish- brownnot observedindistinct????band atperipherywhitish-corneous,with oneL. davini11–14.87.1–10.35–5.51.41–1.670.29angularreddish- brown bandwithout scaly processesindistinct????above andone belowperipherywhitish,L. menorehensis sp. nov.12.4–14.5 (mean 13.4)8.3–10.4 (mean 9.3)5.25–5.75 (mean 5.5)1.31–1.51 (mean 1.44)0.16–0.24 (mean 0.19)roundedwith distinct brown band abovewithout scaly processesindistinctshort, distal part cylindrical, proximal part slightly dilatedabsentlong, broadabsentperiphery
Table 4 (continued). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java.
Table 4 (continued). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java.
Table 4 (continued). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java.