Appendix S1 - Pastoral Practices to Reverse Shrub Encroachment of Sub-Alpine Grasslands: Dung Beetles (Coleoptera, Scarabaeoidea) Respond More Quickly Than Vegetation

Details of the vegetation and dung beetle variables used in GLMM analyses. Table S1, Classification of the species sampled according to four functional diversity traits. The nesting behaviour (N1, dung-ovipositing dweller; N2, soil-ovipositing dweller; N3, tunneller), weight (W1, < 0.005 g; W2, 0.005 g ≤ x ≤ 0.01 g; W3, 0.01 g ≤ x ≤ 0.5 g; W4, > 0.5 g), external protoracical leg length (LL1, < 0.38 mm; LL2, 0.38 mm ≤ x ≤ 0.41 mm; LL3, > 0.41 mm) and tibia width across apex of third tooth and external side (TW1, < 0.125 mm; TW2, 0.125 mm ≤ x ≤0.135 mm; TW3 > 0.135 mm). (DOC)