Nested ANOVA estimates of variance components based on datasets with unequal variances.

Vb is the between-species variance, Vwh the human within-species variance, Veh the human error variance, Vwc the chimpanzee within-species variance and Vec is the chimpanzee error variance. The variance estimates were averaged across 10000 simulations. The true variances used to generate the data are given in brackets. The first set of simulations was based on the average observed variances in humans and chimpanzee, and the chimpanzee error variance and within-species variances were then increased by a factor of 10.