Quantifying the environmental limits to fire spread in grassy ecosystems

Modeling fire spread as an infection process is intuitive: an ignition lights a patch of fuel, which infects its neighbor, and so on. Infection models produce non-linear thresholds, whereby fire spreads only when fuel connectivity and infection probability are sufficiently high. These thresholds are fundamental both to managing fire and to theoretical models of fire spread, whereas applied fire models more often apply quasi-empirical approaches. Here, we resolve this tension by quantifying thresholds in fire spread locally, using field data from individual fires (n=1131) in grassy ecosystems across a precipitation gradient (496-1442mm mean annual precipitation), and evaluating how these scaled regionally (across 533 sites) and across time (1989-2012, 2016-2018) using data from Kruger National Park in South Africa. An infection model captured observed patterns in individual fire spread better than competing models. The proportion of the landscape that burned was well described by measure...