Results of four-way ANOVA (diel * lunar * season * tidal) on the number of click trains per 10 minutes.

The main effects of diel and season, and the interaction effects of diel * lunar, diel * season, diel * tidal, lunar * season, season * tidal, diel * lunar * season, diel * lunar * tidal, diel * season * tidal, lunar * season * tidal and diel * lunar * season * tidal were all significant sources of variability in the occurrence of click trains per 10 minutes. Bold numbers indicate significant effects (P< 0.05).