The genotype and genotype effect of marker and QTL for combining ability and heterosis with two alleles at each locus in BCRIL population.

MM and mm denote the two genotype of molecular marker M; QQ, Qq and qq denote the three genotype of QTL; r represents the recombinant probability between molecular marker M and QTL; μdenote the overall mean value. a and d denote the additive effect and dominant effect, respectively; q and p denote the genotype frequency of QTL QQ and qq in tester, respectively (p+q = 1).*When the genotype of QTL in tester is QQ,, and its genotype frequency is q.#When the genotype of QTL in tester is qq,, and its genotype frequency is p.§When the genotype of QTL in tester is a mixture of QQ and qq, the genotype frequency of QQ and qq are q and p, respectively.