Precision of registration for each barrel column.

The average precision of the soma/dendrites/axon location within the principal column (i.e., containing the neuron's soma) is determined as the standard error of the barrel location (SE). The average precision of long-range projecting axons into columns surrounding the principal column is given by the standard deviation of the barrel location (SD). The minimal precision is derived from the leave-one-out analysis as the root mean squared error between the predicted and actual barrel location (RMSE). The RMSE of the C2 barrel (*) is computed as the average of C1 and C3, because the C2 BC is the origin of the coordinate system during parameterization.