Total abundance of L. harak at each site estimated from an optimal stratified-random sampling design.

Nh represents the theoretical maximum number of transects that could be fit into habitat h without overlap, Wh is the proportional habitat area, nh is the number of transects allocated to habitat h, xh is the mean number of fish per transect for habitat h, sh2 is the variance surrounding the density estimate for habitat h, n is the total number of transects at a site, and s2(xstrat) is the variance surrounding the stratified mean density for the entire site.