Spearman matrix rank correlation coefficients r (999 permutations) between phylogenetic distance matrices MPD (mean pairwise distance) and MNTD (mean nearest neighbor distance and four types of null models: 0–3 [49], [50]1.

1Two different branch length transformation methods were applied with the mean of 999 randomly generated matrices, according to four null models assumptions (see Methods section). ‘All 1’: equal branch lengths assigned to the tree topology; ‘Grafen’s Rho’: Grafen’s branch length transformation method [45]. Significant correlations here indicate that observed sample matrices are not substantially different from random expectations. Null model tests were applied to the three taxa Hesperiidae, Hesperiinae and Pyrginae. Correlations that remain significant after applying a table-wide false discovery rate approach are printed in bold face.Spearman matrix rank correlation coefficients r (999 permutations) between phylogenetic distance matrices MPD (mean pairwise distance) and MNTD (mean nearest neighbor distance and four types of null models: 0–3 [49], [50]1.