Osteology of the Middle Triassic archosaur <i>Lewisuchus admixtus</i> Romer (Chañares Formation, Argentina), its inclusivity, and relationships amongst early dinosauromorphs
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<i>Lewisuchus admixtus</i> is an enigmatic early dinosauriform from the Chañares Formation, Ladinian of Argentina, which has been recently considered a member of Silesauridae. Yet, it differs markedly from Late Triassic silesaurids in dental and vertebral anatomy. Indeed, a detailed redescription of its holotype allowed the identification of several features of the skeleton previously unrecognized amongst silesaurids. These include pterygoid teeth, a dorsomedial posttemporal opening on the otoccipital, foramina associated with cranial nerves X–XII on the caudal region of the prootic–otoccipital, and postaxial neck/trunk vertebrae with craniocaudally expanded neural spines. The presence of a single row of presacral scutes was also confirmed. Some elements previously referred to, or found associated with, the holotype, including a lower jaw, pedal elements and an astragalus, more probably correspond to proterochampsid remains. The anatomical information available for the holotype of <i>L. admixtus</i> was rescored into a new phylogenetic dataset for dinosauromorphs, mostly based on previous works. <i>Lewisuchus admixtus</i> and <i>Pseudolagosuchus major</i> are treated as distinct OTUs because their preserved skeletons mostly lack overlapping parts. The parsimony analysis supports the basal position of <i>L</i>. <i>admixtus</i> within dinosauriforms, prior to the silesaurid–dinosaur split, rather than at the base of Silesauridae. This suggests that a higher number of early dinosauriform clades branched in the Middle and Late Triassic than previously suggested.



