Model parameters.

Most parameters were taken from the original studies that derived the mean field approximations used in the model and are cited accordingly. Other free parameters, like the number of bins of the tonotopic axis N, were fixed to reasonable but arbitrary values at the beginning of the model construction and were not adjusted during the analyses (ad-hoc). Free parameters that were manually tuned are labelled as tuned (x), where x is: 1, for parameters tuned so that the spectral layer integrates the peripheral representation correctly; 2, for parameters tuned to achieve FM-direction selectivity; and 3, for parameters tuned so that the feedback signalling resulted in a fair fit between the model’s pitch predictions and the experimental observations. Short description of the parameters in the last column are further explained along the Methods section. Connectivity parameters are more eloquently described in Fig 13.