Goniopteris ×tico (Thelypteridaceae), a new hybrid fern from Costa Rica

Goniopteris ×tico, a new hybrid fern from La Selva Biological Station in
Heredia Province, Costa Rica, is described based on morphology and
analysis of target-capture DNA sequence data. The hybrid co-occurs with
its two putative progenitors, Goniopteris mollis and Goniopteris
nicaraguensis, and is readily recognizable by its intermediate leaf
dissection and venation. It is also intermediate in pinnae size and shape
and presents irregularly lobed pinnae. Despite the broad overlap in the
geographic distribution of its parental taxa, Goniopteris ×tico is only
known from two collections from a single area of the La Selva Biological
Station, highlighting the importance of close observation of ferns from
even well-collected areas.