Seven favorable WSC alleles individually contributed to significantly higher TGW.

*Significant at P = 0.05; **Significant at P = 0.01; ***Significant at P = 0.001. Total, total WSC at the mid-grain filling produced by leaves and non-leaf organs; Leaf, WSC contributed by current leaf assimilation during the mid-grain filling; Non-leaf, WSC in non-leaf organs at the mid-grain filling, excluding the current leaf assimilation; Remo, WSC used for respiration and remobilization during the mid-grain filling; Ped, peduncle; Low, lower internode; Ste, whole stem; DS, drought stress; WW, well-watered; TGW, 1000-grain weight at maturity.