Estimates (with standard error) of variance components and proportion of variance of anogenital distance in yellow-bellied marmot pups.

AG: additive genetic; MG: maternal genetic; ME: maternal environment; L: litter; Y: year; P: phenotypic; LL: log-likelihood; Variance ratio of AG is the heritability, h2.The raw phenotypic variance of AGD was 13.272. The phenotypic variance of AGD after controlling for the fixed effects was estimated as 6.743 (1.411). Significant random effects are in bold.