A new bird-like dinosaur from the Upper Cretaceous of Mongolia with extremely robust hands supports niche partitioning among velociraptorines

Dromaeosauridae is a clade of bird-like theropod dinosaurs including, among others, the genera Deinonychus and Velociraptor, and characterised by a specialised second toe bearing an enlarged and falciform ungual. Here, we describe an exquisitely-preserved velociraptorine dromaeosaurid from the Upper Cretaceous Djadokhta Formation of Mongolia, and refer it to the new species Shri rapax. This dromaeosaurid is diagnosed by a peculiar combination of vertebral and pelvic features and by an exceptionally robust hand with a very stout pollex bearing the ungual proportionally larger than in any other dromaeosaurid. Combined with cranial adaptations which could support a bite more powerful than in other velociraptorines, the enlarged unguals in both Shri species suggest ecological partitioning in prey preference among the sympatric Djadokhtan dromaeosaurids.