Mean bearings of individual movements throughout the annual cycle.

Movement behavior of sexually immature (0- and 1-year-old) and mature (older than 3-year-old) Lesser Black-backed Gulls (Larus fuscus) at two spatial scales (short movements and long movements). n, sample size; α, mean vector angle; r, mean vector length. Text in bold indicates significance of the mean vector by the Rayleigh test.