Data from: strawberry guava invasion of a Hawaiian rainforest: changing population pattern

# Strawberry guava invasion of a Hawaiian rainforest: Changing population pattern [https://doi.org/10.5061/dryad.dr7sqvb42](https://doi.org/10.5061/dryad.dr7sqvb42) This file provides information on the contents of the file “Psicat Demog 2005-2020 values 20231203.csv”. It is intended to accompany the manuscript titled “Strawberry guava invasion of a Hawaiian rainforest: Changing population pattern” authored by J. S. Denslow, M. T. Johnson, N. L. Chaney, E. C. Farrer. C. C. Horvitz, E. R. Nussbaum, and A. L. Uowolo which appears In the journal *Biotropica*. Please see the “Methods” section of that paper for more detail. The file provides diameter at breast height (DBH, 1.37 m) of vertical strawberry guava (*Psidium cattleyanum* O. Deg. f. *lucidum*) stems measured annually (2005-2020) at 4 study sites in rainforest on windward Hawai'i Island. KAH: Kahauale'a Natural Area Reserve MAK: Puu Maka'ala Natural Area Reserve OLA: Ola'a Forest Reserve WAI: Upper Waiakea Forest Reserve The file provides information on identity of the site, sub-plot identifiers, the tag number of the stem, the type of stem (shoot, sprout), the approximate date of the annual measurement of the site, the diameter of the stem (cm) at breast height (1.37 m) and the status of the stem at the time of measurement. Calculated values for basal area and relative growth rate are also included where available. This information can be used to compare shoots and sprouts with respect to density, stem growth, frequency of leaning or dying, recruitment rate as well as to describe the overall growth and size distribution of the whole population. ## Description of the data and file structure The data file is a spreadsheet with 87 columns and 5847 rows. The first 7 columns provide information on the site, location within the site, tag numbers and stem type. For each measurement date there follow 5 columns providing sampling date, dbh, basal area, relative growth rate, an stem status of a tagged stem. Focal species: *Psidium cattleyanum O. Deg. f* lucidum. (strawberry guava, waiawī) Four replicate plots were established to describe the growth of a strawberry guava population in a lowland *Metrosideros-Cibotium* rainforest on windward Hawaii Island, Hawaii, USA. One plot was established in each reserve. Distances between study sites ranged from 2 to 17 km. See Methods for site descriptions. Site: KAH Kahauale’a Natural Area Reserve; 19o10'N. 155o10'W MAK Pu’u Maka’ala Natural Area Reserve; 19o34'N, 155o11'W OLA Ola’a Forest Reserve, 19o27'N, 155o11'W WAI Upper Waiakea Forest Reserve, 19o35'N, 155o12'W In calculating densities of stems >= 2 cm dbh, the following plot areas apply by site code: KAH Kahauale’a Natural Area Reserve, 0.25 ha plot MAK Pu’u Maka’ala Natural Area Reserve, 0.25 ha plot OLA Ola’a Forest Reserve, 0.15 ha plot WAI Upper Waiakea Forest Reserve, 0.25 ha plot Row, Col: To facilitate annual measurements, each plot was gridded into 5x5m subplots, numbered in the lower left corner in the field. Row and Column coordinates for each subplot are given. SapQuad: All vertical stems >=2 cm dbh were sampled and measured in all subplots. In addition, vertical stems >=0.5 cm to <2cm dbh (saplings) were sampled in a stratified random selection of subplots. In column “SapQuad”, this subsample was coded “1”. The rest of the subplots were coded “0”. In calculating stem densities for each site the following sample areas apply for the sapling subplots: Stems >= 0.5 cm to <2 cm dbh by site KAH 6 subplots, 0.015 ha total MAK 5 subplots, 0.0125 ha OLA 5 subplots, 0.0125 ha WAI 11 subplots, 0.0275 Tag. Tag number given to each measured stem. Numbers are unique within a site but not among sites. NewTag. This number incorporates both a code for the site and the tag number to provide a unique code for each stem in the dataset. StemType. Stems are identified as shoots (SHT: originating from the soil and comprising both stems originating from seed and those originating from the root mat) or sprouts (SPR: originating above ground from an existing larger mother stem). All stems first tagged are vertical (greater than 45 degrees from horizontal). Other leaning and downed stems are also present but not included in this study. A stem subsequently leaning or dying is effectively removed from the data set by recording DBH as “NA”. The following columns are repeated for each measurement date: date2005, date2006, etc. Date of initial or remeasurement as a Julian date. For each site the date reported is the midpoint of the range of dates over which measurements were taken in a given year. dbh2005, dbh2006, etc. Diameter at breast height (1.37m above ground) measured with either calipers (small stems) or a dbh tape in a given year. NA = data not available due to stems not yet recruited or having died or leaned to less than 45 degrees above horizontal since last measured. ba05, ba06, etc. Basal area (cm2) at each year calculated for each stem from its dbh (ba=(dbh/2)2 *). NA = no data. rgr05, rgr06, rgr07, etc. Annual relative growth rate calculated as log (ba at time t+1/ba at time t). Relative growth rate is entered for year t+1 and represents growth over the preceding year. NA = no data. Relative growth rates were not calculated for 2005. test05, test06, etc. A code describing the status of the stem in a particular year as follows: recruitsm = recruit this year into sapling subplot (>=0.5 cm dbh); These stems are considered recruits to the population in subsequent analyses. recruitlg = recruit this year into non-sapling subplot (>= 2 cm dbh); These stems are counted as part of the total population. Stems reaching 2 cm dbh in non-sapling subplots are not coded as recruitlg but are counted in the total population. unborn = stems that recruit in subsequent years but for which we have no prior data notlean = stems vertical (>=45 degrees from the horizontal). This is a place holder. All measured stems are vertical. firstlean = stems leaning to < 45 degrees from horizontal for the first time this year. Leaning stems are deleted from the population by replacing dbh with NA when they first lean and in subsequent years (stilllean) stilllean = stems originally measured as vertical, but subsequently leaning to <= 45 degrees from horizontal. This is a place holder for stems removed from the population by recording NA as dbh for the year. Stems once leaning are assumed to remain below 45 degrees from horizontal. firstdead = stems initially recorded as dead in this year. Dead stems are deleted from the population by replacing dbh with NA when they are first recorded as dead and in subsequent years (stilldead). stilldead = stems dead but originally in the database as live vertical stems NA = none of these categories or missing information