Data from: New information on Titanichthys (Placodermi: Arthrodira) from the Cleveland shale (upper devonian) of Ohio, USA

The placoderm (Titanichthys) from the Late Devonian (Famennian) is based
on incomplete and fragmentary specimens that have hindered understanding
of its overall anatomy and phylogenetic relationships. A new, nearly
complete, articulated specimen from the Upper Devonian Cleveland Shale
provides new information about the previously undescribed rostral,
postmarginal, postsuborbital, submarginal, posterior superognathal plates,
and the nasal capsule. A revised diagnosis is provided for the genus.
Three new diagnostic characters are identified including a transversely
elliptical rostral plate that does not contact adjacent plates, a reduced
posterior superognathal, and a median dorsal plate that inserts into the
posterior dorsal lateral plate. The first comprehensive phylogenetic
analysis of (Titanichthys) indicates that the genus is a basal
aspinothoracid arthrodire closely related to the enigmatic taxa
(Bungartius) and (Tafilalichthys).