TABLE 2 in Formal establishment of Gecarcinoidea H. Milne Edwards, 1837 n. stat. (Crustacea, Decapoda, Brachyura), with recognition of two families, Cardisomatidae n. fam. and Gecarcinidae restrict., and new insights on Epigrapsidae Watabe, 2007

TABLE 2. — Morphological differences between Epigrapsus Heller, 1862 (type species: E. politus) and Grapsodes Heller, 1865 (type species: G. notatus Heller, 1865). Many characters vary considerably with age and size, see Ng et al. 1998; Naruse et al. 2018. Morphological charactersEpigrapsus Heller, 1862 (monospecific: E. politus Heller, 1862)Grapsodes Heller, 1865 (monospecific: G. notatus Heller, 1865)SizesmalllargeCarapacetransversely ovatesquarish to distinctly broader than long, transversely ovate (larger specimens)Dorsal surface of carapacegently convex and swollen, with shallow groovesgently to strongly convex, with rather distinct grooves on large specimensAnterolateral marginarcuate; one very low epibranchial tooth separated from external orbital tooth by very small notch (often undiscernible or, when absent, margin appearing entire)gently convex to strongly arcuate (larger specimens); with three teeth: one external orbital tooth and two epibranchial teeth, variously developed and arranged in variable mannerPosterolateral marginsdistinctly converging towards posterior carapace marginsubparallel to more or less converging towards posterior carapace margin (larger specimens)Antennuleobliquely folded; basal article well developed and exposed; article 2 obliquely folded, entirely visiblehorizontally folded; basal article squarish, exposed; article 2 small, only partially visibleAntennabasal article (2+ 3) J-shaped, bearing article 4 on short inner side; article 5 well visible; short flagellum bent; no orbital hiatusbasal article (2+ 3) squarish; article 4 almost not visible; flagellum bent down in orbital hiatusFrontnot much deflexed, thick, almost entiredeflexed, with inner and outer subfrontal lobes developedOrbitdelimitated, sheltering most part of podophtalmiteonly basally delimitated, sheltering proximal part of podophtalmite; a pointed tooth at inner angleSuborbital ridgethick, tuberculate, possibly acting as pars stridens (inner margin of P1 merus with ridge, possibly acting as plectrum)longer, thinner, smoothProepistomewide, rectangular, with median processsmall, dome-shaped; encircled by developed inner frontal lobesBuccal frameelongate-rectangular, delineated laterallyshorter, quadrate, subcircular to rhomboid, not delineated laterallyEpistome: anterior marginstraight, thick, tuberculateas two concave thin edges joining mediallyMxp3rectangular, distinctly longer than broad, with merus distinctly longer than ischiumshorter, squarish, as broad as long, with merus barely longer than ischiumIschiumwith slight longitudinal sulcus, bordered by setose ridgewith slight short sulcus; no setose ridgeMeruswith very deep oblique depression all along, bordered by setose ridgewith slight depression; no setose ridgePalpinserted near latero-external angle of merus; palp short, with three mobile articles externally visible, first article encroaching on margin of epistomeinserted near middle of anterior margin of merus; palp short, with three mobile articles externally, all included in buccal cavityMxp3 exopodexposed, slender, with flagellumexposed, slender, with vestigial flagellumPterygostomial areanot setosevery setoseSubhepatic areasligthly setosenot setoseSterno-pleonal cavityshallow, its extremity distant from suture 2/3very deep; its extremity not so distant from suture 2/3Male pleonrelatively short; somite 6 wide, with margins straight proximally, convex distallylonger; somite 6 long, narrowing distally, trapezoidal, with straight marginsTelsonshort, bluntly roundedlong, acutely triangularFemale pleonnot describedwide, leaving laterally and especially anteriorly a sternal portion exposed, in addition to episternites; telson in shape of a large triangleThoracic sternumsternite 1 as a spine in continuity with sternite 2; sternite 2 as a large dome-shaped plate, with convex margins; sternites 3 and 4 fused, with slightly sinuous lateral marginssternite 1 as a spine in lower level than sternite 2; sternite 2 narrower, with straight margins; sternites 3 and 4 fused, with slightly concave lateral marginsPleonal locking structure in malesa small button in anterior third of sternite 5; apparatus not functionala largely developed and salient rugose prominence, rather close to suture 4/5. Apparatus seemingly functional: pleonal socket with calcified borderMale gonoporesternal, distant from P5sternal, distant from P5Penisemerging distinctly far from suture 7/8, on middle of sternite 8, crossed by sulcus joining P5 coxo-sternal condyleemerging distinctly far from suture 7/8, on very small sternite 8G1relatively short, stoutrelatively elongated, slenderSetal pouchshort setae on P2 and P3 and on P3 and P4 coxae, forming two small setal pouches between P2/P3 and P3/P4soft and long setae on P3 and P4 coxae forming a large setal pouch between P3/P4