Table 1 in Descriptions of two new species and a new record of the giant pill-millipede genus Zephronia Gray, 1832 from Thailand (Diplopoda, Sphaerotheriida, Zephroniidae)

Table 1 (continued on next six pages). List of species included in this study and their corresponding GenBank accession numbers for the COI gene fragment. SpeciesCOI accession numberVoucher codeLocalityReferenceIngroups1Zephronia chantaburiensis Srisonchai & Wesener, 2024PP754582CUMZ-Zeph0013 (M47)THAILAND, Chanthaburi Province, Tha Mai District, Wat Khao Sukim.Srisonchai et al. (2024)2Zephronia chantaburiensis Srisonchai & Wesener, 2024PP754583CUMZ-Zeph0013 (M48)THAILAND, Chanthaburi Province, Tha Mai District, Wat Khao Sukim.Srisonchai et al. (2024)3Zephronia chrysomallos Bhansali & Wesener, 2022OM509649ZFMK MYR8826THAILAND, Kanchanaburi Province, Sai Yok District, Sai Yok Noi Waterfall.Bhansali & Wesener (2022)4Zephronia dawydoffi Attems, 1953MK330971ZFMK Myr4504N/AWesener (2019)5Zephronia erawani Bhansali & Wesener, 2022OM509650NHMD K56x9THAILAND, Kanchanaburi Province, Si Sawat District, Erawan Waterfall, 50 km W of Kanchanaburi.Bhansali & Wesener (2022)6Zephronia golovatchi Srisonchai, Sutcharit & Likhitrakarn, 2021OM509646ZFMK MYR6262THAILAND, Nakhon Ratchasima Province, Pak Chong District.Bhansali & Wesener (2022)7Zephronia golovatchi Srisonchai, Sutcharit & Likhitrakarn, 2021OM509647ZFMK K53THAILAND, Nakhon Nayok Province, Khao Yai National Park.Bhansali & Wesener (2022)8Zephronia hui Liu & Wesener, 2022OP339790SCAU YGM03CHINA, Guizhou, Tongren City, Jiangkou County, Taiping Town, Yamugou Scenic Area.Zhao et al. (2022)9Zephronia hui Liu & Wesener, 2022OP339791SCAU YGM02CHINA, Guizhou, Tongren City, Jiangkou County, Taiping Town, Yamugou Scenic Area.Zhao et al. (2022)10Zephronia lannaensis Likhitrakarn & Golovatch, 2021OM509629ZFMK MYR3498THAILAND, Chiangmai Province, Mueang District, Doi Suthep.Bhansali & Wesener (2022)11Zephronia lannaensis Likhitrakarn & Golovatch, 2021OM509630ZFMK MYR3501THAILAND, Chiangmai Province, Mae Rim District, Traidhos School Campus.Bhansali & Wesener (2022)12Zephronia lannaensis Likhitrakarn & Golovatch, 2021OM509631ZFMK MYR4911THAILAND, Chiangmai Province, Mae Rim District, Mae Sa Valley.Bhansali & Wesener (2022)13Zephronia lannaensis Likhitrakarn & Golovatch, 2021OM509632NHMD K57BTHAILAND, Chiangmai Province, Mueang District, Doi Suthep, Me Sa Waterfall.Bhansali & Wesener (2022)14Zephronia lannaensis Likhitrakarn & Golovatch, 2021OM509633MHNG 3BTHAILAND, Chiangmai Province, Mueang District, Doi Suthep.Bhansali & Wesener (2022)15Zephronia laotica Wesener, 2019MK330977ZFMK Myr3502LAOS, Champasak Province, east of Mekong, Garden of Erawan Riverside Hotel.Wesener (2019)16Zephronia macula Srisonchai & Wesener, 2024PP754589CUMZ (M54)THAILAND, Sa Kaeo Province, Mueang Sa Kaeo District, Wat Tham Khao Maka.Srisonchai et al. (2024)17Zephronia macula Srisonchai & Wesener, 2024PP754590CUMZ (M55)THAILAND, Sa Kaeo Province, Mueang Sa Kaeo District, Wat Tham Khao Maka.Srisonchai et al. (2024)18Zephronia macula Srisonchai & Wesener, 2024PP754584CUMZ- MYR 0015 (M417)THAILAND, Chanthaburi Province, Khlung District, Thaeo Khlong Khlung Monastery.Srisonchai et al. (2024)19Zephronia macula Srisonchai & Wesener, 2024PP754585CUMZ- MYR 0015 (M418)THAILAND, Chanthaburi Province, Khlung District, Thaeo Khlong Khlung Monastery.Srisonchai et al. (2024)20Zephronia macula Srisonchai & Wesener, 2024PP754586CUMZ- MYR 0015 (M421)THAILAND, Chanthaburi Province, Tha Mai District, Wat Khao Sukim.Srisonchai et al. (2024)21Zephronia macula Srisonchai & Wesener, 2024PP754587CUMZ- MYR 0015 (M422)THAILAND, Chanthaburi Province, Tha Mai District, Wat Khao Sukim.Srisonchai et al. (2024)22Zephronia macula Srisonchai & Wesener, 2024PP754588CUMZ- MYR 0015 (M424)THAILAND, Chanthaburi Province, Tha Mai District, Wat Khao Sukim.Srisonchai et al. (2024)23Zephronia macula Srisonchai & Wesener, 2024PX526095MZKKU (M400)THAILAND, Sa Kaeo Province, Phra Phloeng District, Wat Tham Khao Chan.This study24Zephronia macula Srisonchai & Wesener, 2024PX526096MZKKU (M401)THAILAND, Sa Kaeo Province, Phra Phloeng District, Wat Tham Khao Chan.This study25Zephronia macula Srisonchai & Wesener, 2024PX526097MZKKU (M437)THAILAND, Rayong Province, Khao Chamao District, Wat Tham Suwan Phupha.This study26Zephronia macula Srisonchai & Wesener, 2024PX526098MZKKU (M438)THAILAND, Rayong Province, Khao Chamao District, Wat Tham Suwan Phupha.This study27Zephronia macula Srisonchai & Wesener, 2024PX526099MZKKU (M456)THAILAND, Chachoengsao Province, Phanom Sarakham District, Wat Khao Hin Sorn.This study28Zephronia macula Srisonchai & Wesener, 2024PX526100MZKKU (M457)THAILAND, Chachoengsao Province, Phanom Sarakham District, Wat Khao Hin Sorn.This study29Zephronia macula Srisonchai & Wesener, 2024PX526101MZKKU (M458)THAILAND, Chachoengsao Province, Phanom Sarakham District, Wat Khao Hin Sorn.This study30Zephronia macula Srisonchai & Wesener, 2024PX526102MZKKU (M459)THAILAND, Chachoengsao Province, Phanom Sarakham District, Wat Khao Hin Sorn.This study31Zephronia medongensis Zhao & Liu, 2022OP339793SCAU XZ01CHINA, Xizang Autonomous Region, Medog County.Zhao et al. (2022)32Zephronia ovalis Gray, 1832JX486068ZFMK Myr 0832VIETNAM, Dong Nai Province, Cat Tien National Park.Golovatch et al. (2012)33Zephronia ovalis Gray, 1832PX526103MZKKU (M180)THAILAND, Sisaket Province, Khun Han District, Hui Chan Waterfall.This study34Zephronia ovalis Gray, 1832PX526104MZKKU (M181)THAILAND, Sisaket Province, Khun Han District, Hui Chan Waterfall.This study35Zephronia ovalis Gray, 1832PX526105MZKKU (M204)THAILAND, Surin Province, Bua Chet District, Wat Tham Pha Sai.This study36Zephronia panhai Srisonchai, Sutcharit & Likhitrakarn, 2021OM509643ZFMK MYR 8118THAILAND Ratchaburi Province, Ratchabui-Photharam Districts.Bhansali & Wesener (2022)37Zephronia panhai Srisonchai, Sutcharit & Likhitrakarn, 2021OM509644MHNG 3ATHAILAND, Ratchaburi Province, Chom Bueang District, Tham Kao Bin Forest Park.Bhansali & Wesener (2022)38Zephronia panhai Srisonchai, Sutcharit & Likhitrakarn, 2021OM509645ZFMK MYR 8116THAILAND, Ratchaburi Province, Ratchabui-Photharam Districts.Bhansali & Wesener (2022)39Zephronia phrain Likhitrakarn & Golovatch, 2021OM509634ZFMK MYR 3499THAILAND, Chiang Mai Province, Chiang Dao District, Padeng Lodge.Bhansali & Wesener (2022)40Zephronia phrain Likhitrakarn & Golovatch, 2021OM509635ZFMK MYR3500THAILAND, Chiangmai Province, Mueang District, Doi Suthep.Bhansali & Wesener (2022)41Zephronia phrain Likhitrakarn & Golovatch, 2021OM509636SMFTHAILAND, Chiang Mai Province, Chiang Dao District, Tham Houay Luk.Bhansali & Wesener (2022)42Zephronia phrain Likhitrakarn & Golovatch, 2021OM509637SMFTHAILAND, Chiang Mai Province, Chiang Dao District, Doi Chiang Dao, Ma Lee’s Resort.Bhansali & Wesener (2022)43Zephronia phrain Likhitrakarn & Golovatch, 2021OM509638SMFTHAILAND, Chiang Mai Province, Chai Prakan District, Tham Ngam.Bhansali & Wesener (2022)44Zephronia phrain Likhitrakarn & Golovatch, 2021OM509639ZFMK MYR 4907THAILAND, Chiang Mai Province, Chiang Dao.Bhansali & Wesener (2022)45Zephronia phrain Likhitrakarn & Golovatch, 2021OM509640MHNG 5GTHAILAND, Lamphun Province, Mae Tha District, Doi Khuntan National Park.Bhansali & Wesener (2022)46Zephronia phrain Likhitrakarn & Golovatch, 2021OM509641MHNG 5ITHAILAND, Chiang Mai Province, Chiang Dao District, Doi Chiang Dao.Bhansali & Wesener (2022)47Zephronia phrain Likhitrakarn & Golovatch, 2021OM509642NHMD K35THAILAND, Chiang Mai Province, Ban Musue.Bhansali & Wesener (2022)48Zephronia siamensis Hirst, 1907JX486067.2FMNH-INS-72669THAILAND, Chonburi Province, Sichang District, Koh Sichang.Golovatch et al. (2012)49Zephronia siamensis Hirst, 1907OR530089CUMZTHAILAND, Chonburi Province, Sichang District, Koh Sichang.Srisonchai et al. (2023)50Zephronia siamensis Hirst, 1907PP754591CUMZ (D397)THAILAND, Sa Kaeo Province, Khao Chakan District, Wat Tham Khao Chan.Srisonchai et al. (2024)51Zephronia siamensis Hirst, 1907PP754592CUMZ (M455)THAILAND, Chachoengsao Province, Phanom Sarakham District, Wat Khao Hin Sorn.Srisonchai et al. (2024)52Zephronia sukhothaiensis Srimongkol & Srisonchai, sp. nov.PX526106MZKKU (L42)THAILAND, Sukhothai Province, Ban Dan Lan Hoi District, Wat Khao Fang.This study53Zephronia sukhothaiensis Srimongkol & Srisonchai, sp. nov.PX526107MZKKU (L43)THAILAND, Sukhothai Province, Ban Dan Lan Hoi District, Wat Khao Fang.This study54Zephronia sukhothaiensis Srimongkol & Srisonchai, sp. nov.PX526108MZKKU (M323)THAILAND, Sukhothai Province, Ban Dan Lan Hoi District, Tham Yok Monastery.This study55Zephronia sukhothaiensis Srimongkol & Srisonchai, sp. nov.PX526109MZKKU (M324)THAILAND, Sukhothai Province, Ban Dan Lan Hoi District, Wat Khao Fang.This study56Zephronia sukhothaiensis Srimongkol & Srisonchai, sp. nov.PX526110MZKKU (M325)THAILAND, Sukhothai Province, Ban Dan Lan Hoi District, Tham Yok Monastery.This study57Zephronia sp. (K45 Aow Noi Temple)MW898741NHMD K45THAILAND, Prachuap Kiri Khan Province, Mueang District, Aow Noi Temple.Rosenmejer et al. (2021)58Zephronia sp. (Aow Noi Temple)MW898742ZFMK MYR 8787THAILAND, Prachuap Kiri Khan Province, Mueang District, Aow Noi Temple.Rosenmejer et al. (2021)59Zephronia tratensis Srimongkol & Srisonchai, sp. nov.PX526111MZKKU (M414)THAILAND, Trat Province, Mueang District, Khao Khun Tham Monastery.This study60Zephronia tratensis Srimongkol & Srisonchai, sp. nov.PX526112MZKKU (M415)THAILAND, Trat Province, Mueang District, Khao Khun Tham Monastery.This study61Zephronia tratensis Srimongkol & Srisonchai, sp. nov.PX526113MZKKU (M416)THAILAND, Trat Province, Mueang District, Khao Khun Tham Monastery.This study62Zephronia viridisoma Rosenmejer & Wesener, 2021MW898739NHMD 621695THAILAND, Nakhon Si Thammarat Province, Sichon District, Khao Lark Waterfall.Rosenmejer et al. (2021)63Zephronia viridisoma Rosenmejer & Wesener, 2021MW898740ZFMK MYR 8822THAILAND, Nakhon Si Thammarat Province, Sichon District, Khao Lark Waterfall.Rosenmejer et al. (2021)64Zephronia zhouae Zhao & Liu, 2022OP339794SCAU YN02CHINA, Yunnan Province, Diqing Tibetan Autonomous Prefecture, Laowo Village, Weixi County.Zhao et al. (2022)Outgroups65Sphaerobelum aesculus Rosenmejer & Wesener, 2021MW898737NHMD 621693THAILAND, Phuket Province, Kathu District, Forest.Rosenmejer et al. (2021)66Sphaerobelum cf. aesculus Rosenmejer & Wesener, 2021MW898738NHMD 621694THAILAND, Nakhon Si Thammarat Province, Khao Luang NP.Rosenmejer et al. (2021)67Sphaerobelum benquii Liu & Wesener, 2022OP339792SCAU MMY01CHINA, Guizhou, Tongren City, Jiangkou County, Guanhe Town, Guanhe Village, Maomaoyan.Zhao et al. (2022)68Sphaerobelum bolavensis Wesener, 2019MK330982MHNG LT-10/24LAOS, Champasak Province, Bolaven Plateau, 3 km S of Ban Nong Luang, Tad Kameud.Wesener (2019)69Sphaerobelum denticulatum Wesener, 2019MK330984MHNG LT-10/12LAOS, Oudomxai Province, ca 3 km E of Tad Lak 11, SE of Oudomxai city.Wesener (2019)70Sphaerobelum huzhengkuni Zhao, Yu & Liu, 2020MT657327SCAU SP02CHINA, Guizhou Province, Tongren City, Fanjingshan National Nature Reserve.Zhao et al. (2020)71Sphaerobelum huzhengkuni Zhao, Yu & Liu, 2020MT657328SCAU SP03CHINA, Guizhou Province, Tongren City, Fanjingshan National Nature Reserve.Zhao et al. (2020)72Sphaerobelum lachneeis Wesener, 2019MK330983MHNG LT-10/12LAOS, Oudomxai Province, ca 3 km E of Tad Lak 11, SE of Oudomxai city.Wesener (2019)73Sphaerobelum laoticum Wesener, 2019MK330975SMFLAOS, Vientiane, Province, Vang Vieng.Wesener (2019)74Sphaerobelum meridionalis Bhansali & Wesener, 2022OM509648MHNG 4B-2THAILAND, Yala Province, Bannang Sata District, Bang Lang National Park, near Than To Waterfall.Bhansali & Wesener (2022)75Sphaerobelum nigrum Wesener, 2019MK330976SMFLAOS, Champasak Province, Muang Bachieng, Tad Etu, Ban Lak 35.Wesener (2019)76Sphaerobelum nonghinensis Srikampha & Srisonchai, 2025PQ838982MZKKU (A35)THAILAND, Loei Province, Nong Hin District, Suan Hin Pha Ngam (Hin Pha Ngam Park).Srikampha et al. (2025)77Sphaerobelum nonghinensis Srikampha & Srisonchai, 2025PQ838983MZKKU (A36)THAILAND, Loei Province, Nong Hin District, Suan Hin Pha Ngam (Hin Pha Ngam Park).Srikampha et al. (2025)78Sphaerobelum nonghinensis Srikampha & Srisonchai, 2025PQ838984MZKKU (E28)THAILAND, Loei Province, Nong Hin District, Suan Hin Pha Ngam (Hin Pha Ngam Park).Srikampha et al. (2025)79Sphaerobelum nonghinensis Srikampha & Srisonchai, 2025PQ838985MZKKU (E29)THAILAND, Loei Province, Nong Hin District, Suan Hin Pha Ngam (Hin Pha Ngam Park).Srikampha et al. (2025)80Sphaerobelum nonghinensis Srikampha & Srisonchai, 2025PQ838986MZKKU (M237)THAILAND, Loei Province, Nong Hin District, Suan Hin Pha Ngam (Hin Pha Ngam Park).Srikampha et al. (2025)81Sphaerobelum onyx Srikampha & Srisonchai, 2025PQ838987MZKKU (M211)THAILAND, Buang Kan Province, Seka District, Wat Tham Chum Chang (Nam Yaad Clift).Srikampha et al. (2025)82Sphaerobelum onyx Srikampha & Srisonchai, 2025PQ838988MZKKU (M212)THAILAND, Buang Kan Province, Seka District, Wat Tham Chum Chang (Nam Yaad Clift).Srikampha et al. (2025)83Sphaerobelum peterjaegeri Wesener, 2019MK330972SMF SD553LAOS, Luang Prabang Province, SE Luang Prabang, Nam Khan, Ban Pak Bak, Houay Kho.Wesener (2019)84Sphaerobelum petramurum Srikampha & Srisonchai, 2025PQ838989MZKKU (E5)THAILAND, Khon Kaen Province, Phu Pha Man District, Ban Khao Wong.Srikampha et al. (2025)85Sphaerobelum petramurum Srikampha & Srisonchai, 2025PQ838990MZKKU (L127)THAILAND, Khon Kaen Province, Phu Pha Man District, Ban Khao Wong.Srikampha et al. (2025)86Sphaerobelum petramurum Srikampha & Srisonchai, 2025PQ838991MZKKU (L128)THAILAND, Khon Kaen Province, Phu Pha Man District, Ban Khao Wong.Srikampha et al. (2025)87Sphaerobelum petramurum Srikampha & Srisonchai, 2025PQ838992MZKKU (L6)THAILAND, Khon Kaen Province, Phu Pha Man District, Ban Khao Wong.Srikampha et al. (2025)88Sphaerobelum petramurum Srikampha & Srisonchai, 2025PQ838993MZKKU (M217)THAILAND, Khon Kaen Province, Phu Pha Man District, Ban Khao Wong.Srikampha et al. (2025)89Sphaerobelum petramurum Srikampha & Srisonchai, 2025PQ838994MZKKU (M218)THAILAND, Khon Kaen Province, Phu Pha Man District, Ban Khao Wong.Srikampha et al. (2025)90Sphaerobelum phouloei Wesener, 2019MK330974ZMUC 00040257LAOS, Houaphan Province, Phou Loei.Wesener (2019)91Sphaerobelum schwendingeri Wesener, 2019MK330978MHNG LT-10/03LAOS, Vientiane Province, W. of Vang Vieng, trail to Tham Pou Kham.Wesener (2019)92Sphaerobelum schwendingeri Wesener, 2019MK330981SMFLAOS, Vientiane Province, Vang Vieng, Tham Pou Kham.Wesener (2019)93Sphaerobelum sp. L07MK330979ZMUC 00040261LAOS, Khammouane Province, Ban Khounkham [Khun Kham] (Nahin).Wesener (2019)94Sphaerobelum sp. L10MK330980SMFLAOS, Vientiane Province, Vang Vieng, W. of Nam Song, Tham Nam Or Khem.Wesener (2019)95Sphaerobelum spinatum Wesener, 2019MK330973ZMUC 00040258LAOS, Vientiane Province, Phou Khao Khouay.Wesener (2019)96Sphaerobelum truncatum Wongthamwanich, 2012JN885184FMNH-INS 0000 072 674THAILAND, Nan Province, Song Khwae District, Na Rai Luang Subdistrict, Pang Hi Village.Wongthamwanich et al. (2012)97Sphaerobelum turcosa Srisonchai & Pimvichai, 2023OR530087CUMZ-Zeph0012THAILAND, Loei Province, Mueang Loei District, Phu Pha Lom Forest Park.Srisonchai et al. (2023)98Sphaerobelum turcosa Srisonchai & Pimvichai, 2023OR530087CUMZ-Zeph0012THAILAND, Loei Province, Mueang Loei District, Phu Pha Lom Forest Park.Srisonchai et al. (2023)99Sphaerobelum tujiaphilum Zhao & Liu, 2022OP339783SCAU SD01CHINA, Guizhou, Tongren City, Jiangkou County, Guanhe Town, Sidu Village.Zhao et al. (2022)100Sphaerobelum tujiaphilum Zhao & Liu, 2022OP339784SCAU SD02CHINA, Guizhou, Tongren City, Jiangkou County, Guanhe Town, Sidu Village.Zhao et al. (2022)101Sphaerobelum tujiaphilum Zhao & Liu, 2022OP339785SCAU BHS01CHINA, Guizhou, Tongren City, Jiangkou County, Taiping Town, Baiheshan Village.Zhao et al. (2022)102Sphaerobelum tujiaphilum Zhao & Liu, 2022OP339786SCAU JXT01CHINA, Guizhou, Tongren City, Jiangkou County, Taiping Town, Jiang-xitun Village.Zhao et al. (2022)103Sphaerobelum tujiaphilum Zhao & Liu, 2022OP339787SCAU JXT02CHINA, Guizhou, Tongren City, Jiangkou County, Taiping Town, Jiang-xitun Village.Zhao et al. (2022)104Sphaerobelum tujiaphilum Zhao & Liu, 2022OP339788SCAU SD03CHINA, Guizhou, Tongren City, Jiangkou County, Guanhe Town, Sidu Village.Zhao et al. (2022)105Sphaerobelum tujiaphilum Zhao & Liu, 2022OP339789SCAU DW01CHINA, Guizhou, Tongren City, Jiangkou County, Dewang Town, Xiaobang Village.Zhao et al. (2022)106Arthrosphaera brandtii (Humbert, 1865)FJ409915FMNH-INS 8650TANZANIA, Usambara hills.Wesener et al. (2010)107Cryxus ovalis (Linnaeus, 1758)JX486069.2ZFMK MYR0824VIETNAM, Dong Nai Province, Cat Tien National Park.Golovatch et al. (2012)108Glomeris marginata (Villers, 1789)FJ409909ZFMK Myr009GERMANY, Bonn, Venusberg.Wesener et al. (2010)109Epicyliosoma sp.AF218270NANAEdgecombe & Giribet (2004)110Prionobelum inthanonense Donworth & Wesener, 2024PP297645MHNG 4E-2THAILAND, Chiang Mai Province, Chom Thong District, Doi Inthanon National Park.Donworth & Wesener (2024)111Prionobelum inthanonense Donworth & Wesener, 2024PP297646MHNG 7ATHAILAND, Chiang Mai Province, Chom Thong District, Doi Inthanon National Park.Donworth & Wesener (2024)112Prionobelum naevium Donworth & Wesener, 2024PP297647MHNG 4B-1THAILAND, Yala Province, Than To District, Bang Lang National Park, Than To Waterfall.Donworth & Wesener (2024)113Prionobelum naevium Donworth & Wesener, 2024PP297648NHMD 1184671THAILAND, Yala Province, Than To District, Bang Lang National Park.Donworth & Wesener (2024)114Prionobelum naevium Donworth & Wesener, 2024PP297649NHMD 1184672THAILAND, Yala Province, Than To District, Bang Lang National Park.Donworth & Wesener (2024)115Prionobelum naevium Donworth & Wesener, 2024PP297650NHMD 1184673THAILAND, Yala Province, Than To District, Bang Lang National Park.Donworth & Wesener (2024)116Sphaeromimus splendidus Wesener & Sierwald, 2005FJ409917FMMC-INS 6702MADAGASCAR, Sainte Luce S9Wesener et al. (2010)