Table 2 in Morphology and molecules support the new monotypic genus Parainvolucrella (Rubiaceae) from Asia

Table 2. Morphological comparison of the Hedyotis - Oldenlandia complex distributed in China.
TaxonHabitStipulesFlowersFruitsSeedsPollenDebia Neupane & N. Wikstr.Annual small herbs, erectPapery, broadly triangular, fimbriate with tipped colletersHomostylous with exserted stigma and stamensCompressed globose, loculicidally dehiscentConoidal with deeply depressed exotesta, anticlinal boundaries nearly straight or rounded3-colporate, perforate tectum with psilate muriDimetia (Wight & Arn.) Meisn.Perennial herbs or subshrubs, prostrate, decumbent or climberPapery, truncate, broadly rounded or broadly triangular, spinousHeterostylousSubglobose to ellipsoidal, dehiscent diplophragmously or indehiscentDorsiventrally flattened or trigonous, reticulate, anticlinal boundaries nearly straight3- or 4-colporate, double microreticulate tectum with psilate suprareticulum and microechinate infrareticulumEdrastima Raf.Annual small herbs, decumbentMembranous, truncate, fimbriate with tipped colletersHomostylous with exserted stigma and stamensSubglobose, loculicidally dehiscentTrigonous to ellipsoidal, reticulate, anticlinal boundaries nearly straight3-colporate, microreticulate tectum with psilate muriHedyotis L.Perennial herbs to shrubs, erect or ascendingPapery, triangular, entire to fimbriate with tipped colletersHeterostylous or rarely homostylous with exserted stigma and stamensEllipsoidal, dehiscent diplophragmously or rarely indehiscentDorsiventrally flattened, reticulate, anticlinal boundaries nearly straight3- or 4-colporate, double microreticulate tectum with psilate suprareticulum and microechinate infrareticulumInvolucrella (Hook. f.) Neupane & N. Wikstr.Annual herbs, erect or ascendingPapery, triangular or truncate, margin fimbriate or acicular spinous with tipped colletersHeterostylous or rarely homostylous with included stigma and stamensHemispherical to ellipsoidal, loculicidally dehiscent or indehiscentEllipsoidal, 3-5 pitted, anticlinal boundaries nearly straight or undulate3- or 4-colporate, double microreticulate tectum with psilate suprareticulum and microechinate infrareticulumOldenlandia L.Annual small herbs, erect or ascendingMembranous, flabellate or broadly rounded, fimbriate with tipped colletersHomostylous with included stigma and stamensGlobose to ellipsoidal, loculicidally dehiscentTrigonous, reticulate, anticlinal boundaries nearly straight3- or 4-colporate, microreticulate tectum with psilate muriParainvolucrella R.J. WangAnnual or perennial herbs, decumbentPapery, triangular, fimbriate with tipped colletersHeterostylousSubglobose, 4 longitudinal projections when young, indehiscentTrigonous, reticulate, anticlinal boundaries nearly straight3-colporate, double microreticulate tectum with psilate suprareticulum and microechinate infrareticulumScleromitrion (Wight & Arn.) Meisn.Annual small herbs, erect or ascendingMembranous, triangular to rounded, fimbriate with tipped colletersHomostylous with exserted stigma and stamensSubglobose, loculicidally dehiscentTrigonous to conoidal, reticulate, anticlinal boundaries nearly straight3- or 4-colporate, rugulate tectum with microechinate muriLeptopetalum Hook. & Arn.Annual small herbs, erectPapery, triangular or broadly triangular, fimbriate with tipped colletersHomostylous with included stigma and stamensObconical, winged, loculicidally dehiscentEllipsoidal with deeply depressed exotesta, anticlinal boundaries undulate3-colporate, microreticulate tectum with psilate muri