Frequency (Frq) of occurrence and mean peptide counts of viral pathogens and Nosema in honey bee colonies sampled in 2006, 2007, and 2008a.

aColumns summarize thirty-one colonies from initial CCD study in 2006; eighteen subsamples taken from an observation colony monitored through its collapse from March through August 2007; and a third sample of nine colonies sampled during a CCD incident in Florida in 2008. A hyphen indicates that the value could not be calculated.