Phylogenetic response of naraoiid arthropods to early - middle Cambrian environmental change

The Cambrian Period, primarily known for animal life diversifying, experienced global extinctions. Pulses of extinction in Cambrian Series 2 are exemplified by the disappearance of archaeocyath sponges and olenelline and redlichiid trilobites. However, the effect of such extinctions on outer shelf organisms, as typify Burgess Shale-type (BST) deposits, remains relatively unknown. The phylogeny of naraoiid arthropods, represented in BST deposits globally, has consequently been reconstructed from either side of the Series 2–Miaolingian extinction event to evaluate the response of offshore marine organisms to Cambrian environmental perturbation. As soft anatomy is known for only a subset of naraoiid species, exoskeletal morphology has proven important. Misszhouia and Naraoia (Naraoia) are distinguished morphometrically by posterior shield length/width and anterior shield length/posterior shield length. Morphometry has also been used to strengthen the identification of some cryptic naraoiid...