Multiple factors modulate tree growth complementarity in central European mixed forests

Mixed species forests can often be more productive and deliver higher levels of ecosystem services and functions than monocultures. However, complementarity effects for any given tree species are difficult to generalize because they can vary greatly along gradients of climatic conditions and resource availability. Identifying the conditions where species diversity can positively influence productivity is crucial. To date, few studies have examined how growth complementarity across species and mixture types is modulated by stand and environmental factors, and fewer have considered more than one or two factors. We investigated how complementarity effects for several major central European tree species change with climatic and edaphic conditions, and with stand structural characteristics, including species composition. We used data from the Swiss National Forest Inventory, which is based on 3231 plots of pure and mixed stands (19 mixture types) across a broad environmental gradient, to t...