Analysis of covariance testing whether rates of evolution (<i>S. cerevisiae</i> – <i>S. mikatae</i>) group by occupancy state when region and protein abundance are controlled for.

Data for both all available codons and codons matched across occupancy states (see main text) are shown. L, F, and WP stand for sequence associated with linker, fuzzily and well-positioned nucleosomes, respectively.