PERMANOVA analyses comparing the structure of fish assemblages using multivariate data on the richness and abundance (total <i>max</i> N) of fishes among zone types and among years since the commencement of the zoning plan.

Fish assemblages (a) used Bray-Curtis similarity measures following square root transformation while species richness (b) and total max N (c) used Euclidean distance to generate similarity matrices. Contrasts were included to compare sanctuary zones (SZ) with fished areas (FA) and habitat protection zones (HPZ) with general use zones (GUZ). p-F = pseudo F ratio generated by PERMANOVA.