Estimated variance components (with standard errors in parentheses) for behavioural traits.

The among-focal variance (VF), and residual variance (VR) are shown for all traits along with the among-opponent variance (VO) for traits observed in dyadic trials. Also shown are focal and opponent repeatabilities. P values denote the statistical significance of VF and VO respectively and are derived from 1-tailed likelihood ratio tests (see text for details).
1With models constrained to yield permissible (i.e. non-negative) variance estimates this parameter was bound at the edge of parameter space. Under these conditions standard errors are non-estimable.