Average Spearman rank correlation coefficients across all 50 LODs between all applied measures and fitness.

Since the average number of concepts and their ΣφMax values capture both modular and integrated causal relations in the animat's brain as a whole, they correlated more strongly with fitness than the average number of MC elements, MC concepts and ΦMax>, which nevertheless increased with adaptation. See S1 Fig. for complementary histograms of the correlation coefficients of all individual LODs.Average Spearman rank correlation coefficients across all 50 LODs between all applied measures and fitness.