Table 1 in Thoridae (Crustacea: Decapoda) can penetrate the Abyss: a new species of Lebbeus from the Sea of Okhotsk, representing the deepest record of the family

Table 1 (continued on next 3 pages). Caridean shrimp species hitherto recorded from depths below 3000 m.
Acanthephyridae Spence Bate, 1888Acanthephyra acutifrons Spence Bate, 1888650–4926 mPacificKemp 1906; Chace 1940, 1986; Allen & Butler 1994A. brevirostris Smith, 18851000–5394 mN AtlanticSmith 1885; Domansky 1986A. curtirostris Wood-Mason, 1891190–4970 mPacific, Indian and Atlantic OceansDe Man 1920; Chace 1940, 1986; Crosnier & Forest 1973; Butler 1980; Krygier & Pearcy 1981; Allen & Butler 1994A. eximia Smith, 1884200–5111 mPacific and Atlantic; Mediterranean SeaPohle 1992; Poupin 1996; Linley et al. 2018A. pelagica (Risso, 1816)3635 mSouth OceanBoschi et al. 1981; Wasmer 1986; Iwasaki & Nemoto 1987; Tiefenbacher 1994; Gorny 1999; Basher & Costello 2014A. prionata Foxton, 19711900–4926 mAtlantic, off East Africa; E PacificFoxton 1971; Allen & Butler 1994A. quadrispinosa Kemp, 19395040–5060 mIndian OceanCrosnier 1987A. sica Spence Bate, 1888400–6890 moff New ZealandLörz et al. 2012A. stylorostratis (Spence Bate, 1888)3458 mAtlantic and PacificSpence Bate 1888Acanthephyra spp.6007–6890 mPacificJamieson et al. 2009; Wicksten et al. 2017Heterogenys micro- phthalma (Smith, 1885)3197–5060 mIndian Ocean; NE AtlanticAlcock 1901; Domansky 1986; Crosnier 1987H. monnioti Crosnier, 19872663–4035 mIndian and Atlantic OceansCardoso 2013Hymenodora acanthi- telsonis Wasmer, 19725041–5591 mNW Pacific; NE AtlanticKikuchi & Omori 1985; Domansky 1986H. frontalis Rathbun, 1902586–4432 mN PacificRathbun 1902, 1910; Chace 1986; Marin 2018H. glacialis Buchholz, 1874from near the surface in polar seas to 5610 mNE Atlantic, Pacific and South OceansHavens & Rork 1969; Butler 1980; Just 1980; Wasmer 1986; Domansky 1986; Iwasaki & Nemoto 1987; Hendrickx & Estrada Navarrete 1996; Wicksten 2002; Basher & Costello 2014H. gracilis (Smith, 1886)1000–3733 mSouth OceanSpence Bate 1888; Wasmer 1986; Iwasaki & Nemoto 1987; Gorny 1999; Basher & Costello 2014Meningodora mollis Smith, 1882840–2985 mAtlantic and PacificChace 1940, 1986; Crosnier & Forest 1973; Kensley et al. 1987M. vesca (Smith, 1886)615––5367 mAtlantic and PacificChace 1940, 1986; Crosnier & Forest 1973; Kensley et al. 1987; Allen & Butler 1994Notostomus elegans A. Milne-Edwards, 18810–3500 mAtlantic and PacificCrosnier & Forest 1973; Chace 1986; Kensley et al. 1987N. gibbosus A. Milne-Edwards, 1881850–4000 mAtlantic and PacificCrosnier & Forest 1973; Chace 1986; Kensley et al. 1987Alvinocarididae Christoffersen, 1986Alvinocaris markensis Williams, 19881693–3650 mAtlantic, Mid- Atlantic RidgeWilliams 1988; Wharton et al. 1997; Shank et al. 1999; Komai & Segonzac 2003; Martin & Haney 2005A. methanophila Komai et al., 20052155–3712 mNW AtlanticKomai et al. 2005A. muricola Williams, 19881697–3277 mAtlantic, Gulf of MexicoWilliams 1988; Komai & Segonzac 2005; Komai et al. 2005; Martin & Haney 2005Chorocaris chacei Williams & Rona, 19861600–3650 mAtlantic, Mid- Atlantic RidgeWilliams & Rona 1986; Williams 1987; Martin & Haney 2005; Komai & Segonzac 2008C. vandoverae Martin & Hessler, 19903640 mPacific, Mariana Back Arc BasinMartin & Hessler 1990; Martin & Haney 2005; Nye et al. 2012Mirocaris fortunata (Martin & Christiansen, 1995)850–3480 mAtlantic, Mid- Atlantic RidgeMartin & Christiansen 1995; Vereshchaka 1997; Shank et al. 1999; Komai & Segonzac 2003; Martin & Haney 2005; Komai et al. 2007; Fabri et al. 2011M. indica Komai et al., 20062422–3300 mIndian Ocean, Central Indian RidgeKomai et al. 2006Opaepele susannae Komai et al., 20071500–2986 mAtlantic, Mid- Atlantic RidgeKomai et al. 2007; Beltenev et al. 2009Rimicaris exoculata Williams & Rona, 19861700–4088 mAtlantic, Mid- Atlantic RidgeWilliams & Rona 1986; Martin & Haney 2005; Komai et al. 2007; Komai & Segonzac 2008; Nye et al. 2012R. hybisae Nye et al., 20122300–4960 mAtlantic, CaribbeanNye et al. 2012R. kairei Watabe & Hashimoto, 20022415–3320 mIndian Ocean, Central Ind. RidgeVan Dover et al. 2001; Watabe & Hashimoto 2002; Martin & Haney 2005Bythocarididae Christoffersen, 1987Bythocaris cryonesus Bowman & Manning, 19723803–3805 mArctic: Polar Sea; IcelandBowman & Manning 1972; Just 1980B. curvirostris Kobjakova, 19572352–3965 mArctic: Polar SeaJust 1980Crangonidae Haworth, 1825Neocrangon abyssorum (Rathbun, 1902)887–3200 mN PacificBirštein & Vinogradov 1951; Zarenkov 1965; Hiller-Adams & Case 1985; Kim et al. 2000Parapontophilus abyssi (Smith, 1884)1400–5852 mPacific and AtlanticCrosnier & Forest 1973; Chace 1984; Komai 2008P. occidentalis (Faxon, 1893)837–4082 mE PacificFaxon 1893; Komai 2008; Hendrickx & Papiol 2015P. profundus (Spence Bate, 1888) *4755 mTasman SeaSpence Bate 1888; Komai 2008P. talismani Crosnier & Forest, 19733411–3517 mNE Atlantic; Caribbean SeaCrosnier & Forest 1973; Gore 1985; Komai 2008Placopsicrangon formosa Komai & Chan, 20094807–4824 mW Pacific: TaiwanKomai & Chan 2009Sclerocrangon zenkevitchii Birštein & Vinogradov, 19532995–3950 mN PacificBirštein & Vinogradov 1953; Kim et al. 2000; Komai 2008Disciadidae Rathbun, 1902Lucaya bigelowi Chace, 19394773 mAtlantic: West IndiesChace 1939Eugonatonotidae Chace, 1937Eugonatonotus chacei Chan & Yu, 1991 (as Galatheacaris abyssalis Vereshchaka, 1997) **2000–5000 mW Pacific: SulawesiVereshchaka 1997; Chow et al. 2000Glyphocrangonidae Smith, 1884Glyphocrangon atlantica Chace, 19393400–6373 mAtlanticHolthuis 1971; Gore 1985G. caecescens Wood-Mason in Wood- Mason & Alcock, 18912698–3431 mPacific and IndianKomai 2004; Komai & Komatsu 2016G. longirostris (Smith, 1882)680–3219 mHiller-Adams & Case 1985Nematocarcinidae Smith, 1884Nematocarcinus acanthitelsonis Pequegnat, 19703138–3742 mAtlanticCrosnier & Forest 1973; Gore 1985N. batei Burukovsky, 2000N PacificBurukovsky 2003N. challenger Burukovsky, 20065477 mBurukovsky 2006N. ensifer (Smith, 1882)1000–3600 mAtlanticCrosnier & Forest 1973; Gore 1985N. exilis (Spence Bate, 1888)3300–4000 mAtlantic: Mediterranean SeaCompany et al. 2004N. lanceopes (Spence Bate, 1888)3432 mSouth OceanSpence Bate 1888; Hale 1941; Zarenkov 1968; Gutt et al. 1991, 1994; Gorny 1992, 1999; Arntz et al. 1999; Arntz 2003; Lovrich et al. 2005; Basher & Costello 2014N. longirostris (Spence Bate, 1888)2500–5340 mAtlantic, Pacific, Indian and South OceansSpence Bate 1888; Zarenkov 1968; Iwasaki & Nemoto 1987; Gorny 1999; Burukovsky 2003; Yaldwyn & Webber 2011; Komai & Komatsu 2016N. productus Spence Bate, 1888631–3429 mW PacificChace 1986Oplophoridae Dana, 1852Oplophorus sp.944–5050 mPacific: off New ZealandLörz et al. 2012Systellaspsis braueri (Ba1ss, 1914)200–4000 mN Atlantic and PacificChace 1940, 1986; Crosnier & Forest 1973; Martin & Haney 2005S. cristata (Faxon, 1893)200–3200 mIndian, Pacific and Atlantic OceansHolthuis 1951; Crosnier & Forest 1968; Foxton 1970S. debilis (A. Milne Edwards, 1881)150–5025 mIndian OceanCrosnier 1987; De Man 1920; Chace 1940, 1986; Crosnier & Forest 1973; Baba et al. 1986; Kensley et al. 1987S. pellucida (Filhol, 1884)291–3292 mPacific and AtlanticCrosnier & Forest 1973; Chace 1986; Chan & Yu 1986; Crosnier 1987; Cardoso & Young 2005; Komai et al. 2018Pandalidae Haworth, 1825Stylopandalus richardi (Coutière, 1905)0–3600 mIndian, Pacific and Atlantic OceansChace 1940, 1986; Hayashi & Miyake 1969; Crosnier & Forest 1973; Kensley et al. 1987Pasiphaeidae Dana, 1852Parapasiphae compta Smith, 18844990 mAtlanticCrosnier 1988Pasiphaea scotiae (Stebbing, 1914)3660 mSouth OceanWasmer 1986; Iwasaki & Nemoto 1987; Tiefenbacher 1991, 1994; Gorny 1999; Basher & Costello 2014Sergestidae Dana, 1852Eusergestes arcticus (Krøyer, 1855)3935 mSouth OceanDoflein & Balss 1912; Iwasaki & Nemoto 1987; Tiefenbacher 1994; Gorny 1999Petalidium foliaceum (Spence Bate, 1888)3935 mSouth OceanSpence Bate 1888; Hale 1941; Iwasaki & Nemoto 1987; Tiefenbacher 1991, 1994; Gorny 1999Sergestes arachnipodus (Cocco, 1832)3300–4000 mAtlantic: Mediterranean SeaCompany et al. 2004Sergia robusta (Smith, 1882)3300–4000 mAtlantic: W Ionian SeaCompany et al. 2004Stylodactylidae Spence Bate, 1888Bathylodactylus sp.3922 mPacific: Clarion- Clipperton ZoneAmon et al. 2017Bathystylodactylus bathyalis (Cleva, 1994)3502–3515 mW Pacific: Coral SeaCleva 1994B. cf. bathyalis (Cleva, 1994)4826 mPacific: Marianas Trench Marine Natl MonumentWicksten et al. 2017B. inflatus Hanamura & Takeda, 19963436–3452 mW Pacific: TaiwanHanamura & Takeda 1996Thoridae Kingsley, 1879Lebbeus sokhobio sp. nov.3301–3366 mNW Pacific: Sea of OkhotskMarin 2018; present study
* Known from single holotype specimen.
** G. abyssalis is a mesopelagic larva of E. chacei. The benthic species itself has not been recorded deeper than 1000 m, but G. abyssalis can be collected in mid-water when trawls are hauled up from abyssal depths. Nevertheless, the records of G. abyssalis and E. chacei, respectively, are presented here as they were published in the literature.