PC parameters describing the two-compartment rate model.

, and denote the total strength of connection between PV neurons and the soma of PCs, SOM neurons and the dendrites of PCs and the recurrence strength between PCs, respectively. The total connection strength is given by the product of the number of existing connections between two neuron types (or compartments) and the strength for individual connections. All parameters taken from Murayama et al. (2009) [34]. Note that we incorporated the gain factor present in Murayama et al. (2009) [34] into the parameters to achieve unit consistency for all neuron types.