Table 1 in Description of two new species of Desmoscolex Claparède, 1863 (Nematoda: Desmoscolecidae) from the Cassidaigne Canyon, Mediterranean Sea

Table 1 (continued on next four pages). List of valid species within the genus Desmoscolex Claparède, 1863 including their respective authors, their synonyms and the environments documented in their original descriptions. Species marked with * are synonymized within the corresponding subgenus for the first time in this study, based on the original description of both the subgenus and the species. valid specie sauthorenvironmentsynonymized namesDesmoscolex (Desmoscolex) abyssorum*Decraemer, 1984marineDesmoscolex abyssorum Decraemer, 1984Desmoscolex (Desmoscolex) adenotrichus*Lorenzen, 1969marineDesmoscolex adenotrichus Lorenzen, 1969Desmoscolex (Desmoscolex) adriaticus*Schepotieff, 1907marineDesmoscolex adriaticus Schepotieff, 1907Desmoscolex (Desmoscolex) algivorusCoomans, Vincx & Decraemer, 1985fresh waterDesmoscolex algivorus Coomans, Vincx & Decraemer, 1985Desmoscolex (Desmoscolex) amaurus*Lorenzen, 1972marineDesmoscolex amaurus Lorenzen, 1972Desmoscolex (Desmoscolex) americanus*Chitwood, 1936marineDesmoscolex americanus Chitwood, 1936Desmoscolex (Desmoscolex) annulatus*Schepotieff, 1907marineDesmoscolex annulatus Schepotieff, 1907Desmoscolex (Desmoscolex) antarcticos(Timm, 1970)marineDesmoscolex antarcticos (Timm, 1970); Protodesmoscolex antarcticos Timm, 1970Desmoscolex (Desmoscolex) aquaedulcisStammer, 1935fresh waterDesmoscolex aquaedulcis Stammer, 1935Desmoscolex (Desmoscolex) articulatus*Timm, 1978marineDesmoscolex articulatus Timm, 1978Desmoscolex (Desmoscolex) asetosus*Decraemer, 1975marineDesmoscolex asetosus Decraemer, 1975Desmoscolex (Desmoscolex) australicus*Decraemer, 1974marineDesmoscolex australicus Decraemer, 1974Desmoscolex (Desmoscolex) balticus*Lorenzen, 1971marineDesmoscolex balticus Lorenzen, 1971Desmoscolex (Desmoscolex) bathyalis*Freudenhammer, 1975marineDesmoscolex bathyalis Freudenhammer, 1975Desmoscolex (Desmoscolex) bathybiusTimm, 1970marineDesmoscolex bathybius Timm, 1970Desmoscolex (Desmoscolex) bengalensis*Timm, 1961marineDesmoscolex bengalensis Timm, 1961Desmoscolex (Desmoscolex) borealis*Kreis, 1963marineDesmoscolex borealis Kreis, 1963Desmoscolex (Desmoscolex) brachyrhynchus*Bussau, 1993marineDesmoscolex brachyrhynchus Bussau, 1993Desmoscolex (Desmoscolex) brevisetosus*Decraemer, 1974marineDesmoscolex brevisetosus Decraemer, 1974Desmoscolex (Desmoscolex) californicusTimm, 1970marine; brackishDesmoscolex californicus Timm, 1970Desmoscolex (Desmoscolex) campbelli*Allgén, 1946marineDesmoscolex campbelli Allgén, 1946Desmoscolex (Desmoscolex) chaetalatus*Freudenhammer, 1975marineDesmoscolex chaetalatus Freudenhammer, 1975Desmoscolex (Desmoscolex) chaetogaster*Greeff, 1869marineDesmoscolex chaetogaster Greeff, 1869; Eudesmoscolex chaetogaster (Greeff, 1869) Schuurmans stekhoven 1935Desmoscolex (Desmoscolex) complexus*Decraemer, 1984marineDesmoscolex complexus Decraemer, 1984Desmoscolex (Desmoscolex) conurus*Steiner, 1916marineDesmoscolex conurus Steiner, 1916Desmoscolex (Desmoscolex) coronatus*Soetaert, 1989marineDesmoscolex coronatus Soetaert, 1989Desmoscolex (Desmoscolex) cosmopolitesTimm, 1970marineDesmoscolex cosmopolites Timm, 1970Desmoscolex (Desmoscolex) cristatus*(Allgén, 1932) Lorenzen, 1971marineDesmoscolex cristatus (Allgén, 1932) Lorenzen 1971; Eudesmoscolex cristatus (Allgén, 1932)valid specie sauthor environmentsynonymized namesDesmoscolex (Desmoscolex) curvespiculatum*Decraemer, 1984marineDesmoscolex curvespiculatum Decraemer, 1984Desmoscolex (Desmoscolex) deconincki*Decraemer, 1974marineDesmoscolex deconincki Decraemer, 1974Desmoscolex (Desmoscolex) decraemerae*Soetaert, 1989marineDesmoscolex decraemerae Soetaert, 1989Desmoscolex (Desmoscolex) demerarae*Decraemer, 1983marineDesmoscolex demerarae Decraemer, 1983Desmoscolex (Desmoscolex) dimorphus*Decraemer, 1974marineDesmoscolex dimorphus Decraemer, 1974Desmoscolex (Desmoscolex) draconemoidesTimm, 1970marineDesmoscolex draconemoides Timm, 1970Desmoscolex (Desmoscolex) dussartiJuget, 1969fresh waterDesmoscolex dussarti Juget, 1969Desmoscolex (Desmoscolex) eftus*Freudenhammer, 1975marineDesmoscolex eftus Freudenhammer, 1975Desmoscolex (Desmoscolex) falcatus*Lorenzen, 1972marineDesmoscolex falcatus Lorenzen, 1972Desmoscolex (Desmoscolex) fennicus*Lorenzen, 1969marineDesmoscolex fennicus Lorenzen, 1969Desmoscolex (Desmoscolex) frigidus*Timm, 1978marineDesmoscolex frigidus Timm, 1978Desmoscolex (Desmoscolex) galeatus*Freudenhammer, 1975marineDesmoscolex galeatus Freudenhammer, 1975Desmoscolex (Desmoscolex) geraerti*Decraemer, 1974marineDesmoscolex geraerti Decraemer, 1974Desmoscolex (Desmoscolex) gerlachiTimm, 1970marineDesmoscolex gerlachi Timm, 1970Desmoscolex (Desmoscolex) gladisetosusTimm, 1970marineDesmoscolex gladisetosus Timm, 1970Desmoscolex (Desmoscolex) grandiamphis*Inglis, 1967marineDesmoscolex grandiamphis Inglis, 1967Desmoscolex (Desmoscolex) granulatus*Decraemer, 1975marineDesmoscolex granulatus Decraemer, 1975Desmoscolex (Desmoscolex) italicusTimm, 1970marineDesmoscolex italicus Timm, 1970Desmoscolex (Desmoscolex) keiensis*Kreis, 1934marineDesmoscolex keiensis Kreis, 1934Desmoscolex (Desmoscolex) koloensisDecraemer, 1984fresh water; terrestrialDesmoscolex koloensis Decraemer, 1984Desmoscolex (Desmoscolex) labiosus*Lorenzen, 1969marineDesmoscolex labiosus Lorenzen, 1969Desmoscolex (Desmoscolex) laevis*Kreis, 1928marineDesmoscolex laevis Kreis, 1928Desmoscolex (Desmoscolex) lanceosetatusJung, Kihm & Rho, 2024marineDesmoscolex lanceosetatus Jung, Kihm & Rho, 2024Desmoscolex (Desmoscolex) lanuginosus *Panceri, 1876marineDesmoscolex lanuginosus Panceri, 1876Desmoscolex (Desmoscolex) lapilliferus*Freudenhammer, 1975marineDesmoscolex lapilliferus Freudenhammer, 1975Desmoscolex (Desmoscolex) lemaniJuget, 1969fresh waterDesmoscolex lemani Juget, 1969Desmoscolex (Desmoscolex) leptus*Steiner, 1916marineDesmoscolex leptus Steiner, 1916Desmoscolex (Desmoscolex) lissus*Steiner, 1916marineDesmoscolex lissus Steiner, 1916Desmoscolex (Desmoscolex) longiamphisTimm, 1970marineDesmoscolex longiamphis Timm, 1970Desmoscolex (Desmoscolex) longirostris*Bussau, 1993marineDesmoscolex longirostris Bussau, 1993Desmoscolex (Desmoscolex) longisetosusTimm, 1970marineDesmoscolex longisetosus Timm, 1970Desmoscolex (Desmoscolex) lorenzeni*Freudenhammer, 1975marineDesmoscolex lorenzeni Freudenhammer, 1975Desmoscolex (Desmoscolex) macramphis*Decraemer, 1984marineDesmoscolex macramphis Decraemer, 1984valid specie sauthor environment synonymized namesDesmoscolex (Desmoscolex) macrophasmatus *Decraemer, 1984marineDesmoscolex macrophasmatus Decraemer, 1984; Desmoscolex macrophasmata Decraemer, 1984Desmoscolex (Desmoscolex) maxTimm, 1970marineDesmoscolex max Timm, 1970Desmoscolex (Desmoscolex) membranifer*Soetaert, 1989marineDesmoscolex membranifer Soetaert, 1989Desmoscolex (Desmoscolex) membranosus*Decraemer, 1974marineDesmoscolex membranosus Decraemer, 1974Desmoscolex (Desmoscolex) michaelseni*Steiner, 1916marineDesmoscolex michaelseni Steiner, 1916Desmoscolex (Desmoscolex) minor*Schepotieff, 1907marineDesmoscolex minor Schepotieff, 1907Desmoscolex (Desmoscolex) minutus*Claparède, 1863marineDesmoscolex minutus Claparède, 1863Desmoscolex (Desmoscolex) multiannulatus*Decraemer, 1983marineDesmoscolex multiannulatus Decraemer, 1983Desmoscolex (Desmoscolex) nanus*Steiner, 1916marineDesmoscolex nanus Steiner, 1916Desmoscolex (Desmoscolex) nibelungussp. nov.marineDesmoscolex (Desmoscolex) nudus*Chitwood, 1951marineDesmoscolex nudus Chitwood, 1951Desmoscolex (Desmoscolex) nymphianus*Decraemer, 1974marineDesmoscolex nymphianus Decraemer, 1974Desmoscolex (Desmoscolex) obscurus*Bussau, 1993marineDesmoscolex obscurus Bussau, 1993Desmoscolex (Desmoscolex) opacus*Bussau, 1993marineDesmoscolex opacus Bussau, 1993Desmoscolex (Desmoscolex) parabyssorum*Soetaert, 1989marineDesmoscolex parabyssorum Soetaert, 1989Desmoscolex (Desmoscolex) parafalklandiae* Allgén, 1955marineDesmoscolex parafalklandiae Allgén, 1955Desmoscolex (Desmoscolex) parakoloensisDecraemer, 1984fresh waterDesmoscolex parakoloensis Decraemer, 1984Desmoscolex (Desmoscolex) paraleptus*Decraemer, 1975marineDesmoscolex paraleptus Decraemer, 1975Desmoscolex (Desmoscolex) paralongisetosus *Decraemer, 1983marineDesmoscolex paralongisetosus Decraemer, 1983Desmoscolex (Desmoscolex) parvospiculatus* Decraemer, 1996marineDesmoscolex parvospiculatus Decraemer, 1996Desmoscolex (Desmoscolex) pelophilus*Steiner, 1916marineDesmoscolex pelophilus Steiner, 1916Desmoscolex (Desmoscolex) perspicuus*Freudenhammer, 1975marineDesmoscolex perspicuus Freudenhammer, 1975Desmoscolex (Desmoscolex) petalodes*Lorenzen, 1972marineDesmoscolex petalodes Lorenzen, 1972Desmoscolex (Desmoscolex) prampramensis*Steiner, 1916marineDesmoscolex prampramensis Steiner, 1916Desmoscolex (Desmoscolex) proboscis *Lorenzen, 1972marineDesmoscolex proboscis Lorenzen, 1972Desmoscolex (Desmoscolex) pusillus*Lorenzen, 1969marineDesmoscolex pusillus Lorenzen, 1969Desmoscolex (Desmoscolex) pustulatus*Freudenhammer, 1975marineDesmoscolex pustulatus Freudenhammer, 1975Desmoscolex (Desmoscolex) quadricomoidesTimm, 1970marineDesmoscolex quadricomoides Timm, 1970Desmoscolex (Desmoscolex) remiferTimm, 1970marineDesmoscolex remifer Timm, 1970Desmoscolex (Desmoscolex) roscoffiensis*Decraemer, 1979marineDesmoscolex roscoffiensis Decraemer, 1979Desmoscolex (Desmoscolex) rostratusTimm, 1970marineDesmoscolex rostratus Timm, 1970Desmoscolex (Desmoscolex) rotundicephalusJung, Kihm & Rho, 2024marineDesmoscolex rotundicephalus Jung, Kihm & Rho, 2024Desmoscolex (Desmoscolex) rudolphi *Steiner, 1916marineDesmoscolex rudolphi Steiner, 1916valid specie sauthor environment synonymized namesDesmoscolex (Desmoscolex) segonzaci*Decraemer, 1983marineDesmoscolex segonzaci Decraemer, 1983Desmoscolex (Desmoscolex) sieverti*Freudenhammer, 1975marineDesmoscolex sieverti Freudenhammer, 1975Desmoscolex (Desmoscolex) spinirostris*Decraemer, 1984marineDesmoscolex spinirostris Decraemer, 1984Desmoscolex (Desmoscolex) spinosus*Decraemer, 1976marineDesmoscolex spinosus Decraemer, 1976Desmoscolex (Desmoscolex) tenuiseta*Filipjev, 1922marineDesmoscolex tenuiseta Filipjev, 1922Desmoscolex (Desmoscolex) timmi*Decraemer, 1974marineDesmoscolex timmi Decraemer, 1974Desmoscolex (Desmoscolex) valkyriasp. nov.marineDesmoscolex (Desmoscolex) vanoyei*De Coninck, 1943marineDesmoscolex vanoyei De Coninck, 1943Desmoscolex (Desmoscolex) variabilis*Soetaert, 1989marineDesmoscolex variabilis Soetaert, 1989Desmoscolex (Desmoscolex) veliferTimm, 1970marineDesmoscolex velifer Timm, 1970Desmoscolex (Desmoscolex) vinealis*Weischer, 1962terrestrialDesmoscolex vinealis Weischer, 1962Desmoscolex (Desmoscolex) yongei*Decraemer, 1974marineDesmoscolex yongei Decraemer, 1974Desmoscolex (Desmolorenzenia) camerunensisDecraemer & Sturhan, 1997terrestrial–Desmoscolex (Desmolorenzenia) cooleni(Decraemer, 1978)marineDesmoscolex cooleni (Decraemer, 1978); Desmolorenzenia cooleni Decraemer, 1978Desmoscolex (Desmolorenzenia) coreensisRho, Kim & Chang, 2007marineDesmoscolex coreensis Rho, Kim & Chang, 2007Desmoscolex (Desmolorenzenia) crassicaudus (Timm, 1970)marineDesmoscolex crassicauda (Timm, 1970); Desmolorenzenia crassicauda (Timm, 1970)Desmoscolex (Desmolorenzenia) curvicaudus(Timm, 1978)marineQuadricoma curvicauda Timm, 1978; Tricoma (Quadricoma) curvicauda (Timm, 1978)Desmoscolex (Desmolorenzenia) desmoscolecoides(Timm, 1970)marineDesmolorenzenia desmoscolecoides (Timm, 1970); Desmoscolex desmoscolecoides (Timm, 1970); Quadricoma desmoscolecoides Timm, 1970; Tricoma (Quadricoma) desmoscolecoides Timm, 1970Desmoscolex (Desmolorenzenia) eurycricusFilipjev, 1922marineDesmolorenzenia eurycricus (Filipjev, 1922); Desmolorenzenia longicauda (Timm, 1970); Desmoscolex (Desmolorenzenia) longicauda Timm, 1970; Desmoscolex eurycricus Filipjev, 1922; Desmoscolex longicauda (Timm, 1970); Quadricoma eurycricus (Filipjev, 1922); Quadricoma longicauda Timm, 1970; Tricoma (Quadricoma) eurycricus Filipjev, 1922; Tricoma (Quadricoma) longicauda (Timm, 1970)Desmoscolex (Desmolorenzenia) frontalisGerlach, 1952marineDesmoscolex frontalis Gerlach, 1952Desmoscolex (Desmolorenzenia) gourbaultaeDecraemer, 1986marineDesmoscolex gourbaultae Decraemer, 1986Desmoscolex (Desmolorenzenia) hupferi(Steiner, 1916)marineDesmoscolex hupferi Steiner, 1916; Desmolorenzenia hupferi (Steiner, 1916); Quadricoma hupferi (Steiner, 1916) Timm, 1970; Tricoma (Quadricoma) hupferi (Steiner, 1916)Desmoscolex (Desmolorenzenia) montanus(Decraemer & Sturhan, 1982)terrestrialDesmolorenzenia montana Decraemer & Sturhan, 1982; Desmolorenzenia montanus Decraemer & Sturhan, 1982; Desmoscolex (Desmolorenzenia) montana (Decraemer & Sturhan, 1982); Desmoscolex montana (Decraemer & Sturhan, 1982) Decraemer 1984Desmoscolex (Desmolorenzenia) parvus(Timm,1970)marineDesmolorenzenia parva (Timm, 1970); Desmoscolex parvus (Timm, 1970); Quadricoma parva Timm, 1970; Tricoma (Quadricoma) parva Timm, 1970Desmoscolex (Desmolorenzenia) pedunculusRho, Kim & Chang, 2007marineDesmoscolex pedunculus Rho, Kim & Chang, 2007Desmoscolex (Desmolorenzenia) platycricusSteiner, 1916marineDesmoscolex platycricus Steiner, 1916; Desmolorenzenia platycricus Steiner, 1916Desmoscolex (Desmolorenzenia) vittatusLorenzen, 1969marineDesmolorenzenia vittata (Lorenzen, 1969); Desmoscolex vittatus Lorenzen, 1969; Quadricuma vittata (Lorenzen, 1969); Tricoma (Quadricoma) vittata (Lorenzen, 1969)Desmoscolex (Pareudesmoscolex) laciniosusLorenzen, 1969marine; brackishPareudesmoscolex laciniosus Lorenzen, 1969Desmoscolex (Pareudesmoscolex) lacustrisJuget, 1969fresh waterPareudesmoscolex lacustris Juget, 1969Desmoscolex (Pareudesmoscolex) papillosus(Schulz, 1935)marine; brackishEudesmoscolex papillosus Schulz, 1935; Pareudesmoscolex papillosus (Schulz, 1935)Desmoscolex (Pareudesmoscolex) pratensisLorenzen, 1969marinePareudesmoscolex pratensis Lorenzen, 1969Desmoscolex (Protricomoides) pravusDecraemer, 1985marine–Desmoscolex (Protricomoides) squamosusTimm, 1970marineDesmoscolex squamosus (Timm, 1970)Desmoscolex (Protricomoides) noctuabundus*Bussau, 1993marineDesmoscolex noctuabundus Bussau, 1993