Multitrophic diversity and biotic associations influence subalpine forest ecosystem multifunctionality

Biodiversity across multiple trophic levels is required to maintain multiple ecosystem functions. Yet, it remains unclear how multitrophic diversity and species interactions regulate ecosystem multifunctionality. Here, combining data from nine different trophic groups (including trees, shrubs, herbs, leaf mites, small mammals, bacteria, pathogenic fungi, saprophytic fungi and symbiotic fungi) and 13 ecosystem functions related to supporting, provisioning and regulating services, we used a multitrophic perspective to evaluate the effects of elevation, diversity and network complexity on scale-dependent subalpine forest multifunctionality. Our results demonstrate that elevation and soil pH significantly modified species composition and richness across multitrophic groups and influenced multiple functions simultaneously. We provide evidence that species richness across multiple trophic groups had stronger effects on multifunctionality than species richness at any single trophic level. More...