Table 1 in Checklist of Water mites in Mexico. Historical background and DNA barcoding perspectives

Table 1 Checklist of water mite species in Mexico. * Without data collection.
SpeciesStateHabitatReferenceArrenuridaeArrenurus (Arrenurus) dentipetiolatusOaxaca, GuanajuatoPondMarshall 1908Arrenurus (Arrenurus) valenciusCampeche, TabascoWater-filled roadsideMarshall 1919Arrenurus (Arrenurus) munovusChiapasStreamCook 1980Arrenurus (Arrenurus) wucabusOaxacaPondCook 1980Arrenurus (Arrenurus) tamauulipensisTamaulipasLakeCook 1980Arrenurus (Arrenurus) xochimilcoensisMexico CityStanding watersCramer and Cook 1992Arrenurus (Megaluracarus) manubriatorTabascoPondMarshall 1903Arrenurus (Megaluracarus) birgeiCampecheLagoonMarshall 1903Arrenurus (Megaluracarus) marshallaeCampecheWater-filled ditchPiersig 1904Arrenurus (Megaluracarus) gricalusVeracruzPondCook 1980Arrenurus (Megaluracarus) hartesusTabascoPondCook 1980Arrenurus (Megaluracarus) neoexpansusTabascoPondCook 1980Arrenurus (Megaluracarus) tabascoensisCampecheWater-filled ditchCook 1980Arrenurus (Megaluracarus) trassamusTabascoPondCook 1980Arrenurus (Megaluracarus) zitavusCampecheWater filled-ditchCook 1980Arrenurus (Megaluracarus) campechensisCampecheWater filled-ditchCook 1980Arrenurus (Megaluracarus) wolardusVeracruz, ColimaPondCook 1980Arrenurus (Megaluracarus) costeroaeTamaulipasLakeCramer and Cook 1992Arrenurus (Megaluracarus) alloexpansusColimaNot specifiedCramer and Cook 1992Arrenurus (Megaluracarus) apizanusTamaulipasLakeCramer and Cook 1992Arrenurus (Megaluracarus) catoiTamaulipasLakeCramer and Cook 1992Arrenurus (Megaluracarus) champayanusTamaulipasLakeCramer and Cook 1992Arrenurus (Megaluracarus) colitusTamaulipasLakeCramer and Cook 1992Arrenurus (Megaluracarus) anaeTamaulipasLakeCramer and Cook 1998Arrenurus (Megaluracarus) anitahoffmannaeTabascoLake, pond, canalRamirez-Sánchez and Rivas 2013Arrenurus (Megaluracarus) olmecaTabascoLake, pond, canalRamirez-Sánchez and Rivas 2013Arrenurus (Megaluracarus) mayaYucatán, Quintana RooCenoteRamirez-Sánchez and Rivas 2013Arrenurus (Megaluracarus) urbanusMexico cityCanalRamirez-Sánchez and Rivas 2013Arrenurus (Megaluracarus) eduardoiQuintana RooPool (in a stream)Montes-Ortiz et al.2022Arrenurus (Megaluracarus) federicoiQuintana RooPool (in a stream)Montes-Ortiz et al.2022Arrenurus (Megaluracarus) ecosurQuintana RooCenote, lagoon, wetlandsMontes-Ortiz et al.2022Arrenurus (Megaluracarus) beatrizaeQuintana Roo, TabascoWetland, lagoonMontes-Ortiz et al.2022Arrenurus (Dadayella) zempoalaMexico stateSmall streamCook 1980Arrenurus (Dadayella) adrianaeColima, MichoacanWetland, lagoonCramer and Cook 1992Arrenurus (Dadayella) veracruzensisVeracruzPondCramer and Cook 1992Arrenurus (Dadayella) aztecusVeracruzWetland, lagoonCramer and Cook 1992Arrenurus (Dadayella) colimensisColimaWetland, lagoonCramer and Cook 1992Arrenurus (Dadayella) cristinaeQuintana RooWetlandMontes-Ortiz et al.2022Arrenurus (Truncaturus) plevamusGuerreroSmall streamCook 1980Arrenurus (Truncaturus) zukovusChiapasGravel-bottom streamCook 1980Arrenurus (Truncaturus)teoceloensisVeracruzStreamRivas and Cramer 1992Arrenurus (?Arrhenuropsis) mexicanusTamaulipas, ColimaLagoonCramer and Cook 1992Arrenurus (?) nayaritensisNayaritSmall streamCook 1980HydrachnidaeHydrachna miliariaOaxacaPondBerlese 1888 in Cook 1980Hydrachna leovazquezaeMexico stateTemporal pondCramer and Costero 1986Hydrachna guanajuatensisGuanajuatoPondCook 1980Hydrachna mexicanaVeracruzPondCook 1980Hydrachna rotundaOaxaca, Nayarit, Guanajuato.PondMarshall 1930 in Cook 1980LimnocharidaeRhyncholimnochares (Rhyncholimnochares)ChiapasRiverCook 1980Rhyncholimnochares (Paralimnochares)ChiapasRiverCook 1980EylaidaeEylais guanajuatensisGuanajuato*Dugés 1873Eylais mexicanaMexico stateHelocrene springCook 1980HydriyphantidaeHydryphantes (Hydryphantes) ramosus ramosus OaxacaPondDaday 1905 in Cook 1980Thyopsella obscuraMexico state.Helocrene springCook 1980ProtziaQuerétaroStreamGoldschmidt et al. 2015HydrodromidaeHydrodroma peregrina perigrinaCampeche, GuanajuatoPondLundblad 1941 in Cook 1980Hydrodroma (Clavipes) despisciensOaxaca, Chiapas, Veraruz, San Luis Potosí.Stream, river.Lundblad 1953 in Cook 1980RhynchohydracaridaeClathrosperchon punctatusOaxaca, Chiapas, Veracruz.Matted roots, bottom of streamCook 1980SperchontidaeSperchon mexicanusMexico stateStreamCook 1980Sperchon neotropicusOaxaca, Coahuila, Veracruz.Algae on rocks, streamCook 1980Sperchon (Hispidopershon) gledhilliOaxaca, Veracruz, Guerrero.StreamViets 1977Sperchon (Acadiosperchon) pontifexOaxacaSubmerged vegetationOtero-Colina 1987aSperchon (Mixosperchon) oaxacensisOaxacaSubmerged vegetationOtero-Colina 1987aAnisitsiellidaeBandakia hoffmannaeMexico state.SpringCramer and Smith 1991Bandakia mexicanaMexico state.SpringCramer and Smith 1991Mamersella mesoamericanaTabascoLagoonOtero-Colina 1987aMamersellides costeniusVeracruz, Colima, Estados Unidos (Florida)Ponds, Marshes, backwater pools of streams.Cramer and Smith 1993LebertiidaeLebertia (Pseudolebertia) aztecaMexico stateStreamCook 1980OxidaeOxus dugesiVeracruz, Costa RicaStreamCook 1980Oxus stolliGuerreroTributary riverCook 1980TorrenticolidaeTorrenticola (Monoatractides) lembadaChiapas, Costa RicaStream, river.Cook 1980Torrenticola (Monoatractides) gordaNayarit, Guerrero, OaxacaRiverCook 1980Torrenticola (Monoatractides) veracruzensisVeracruz, ChiapasStreamCook 1980Torrenticola (Torrenticola) ralaGuerreroRiverCook 1980Torrenticola (Torrenticola) keesdavidsiMexico state.StreamCramer 1992Torrenticola (Torrenticola) carlbaderiMexico state.StreamCramer 1992Torrenticola (Torrenticola) esbeltaMexico state.StreamCramer 1992Torrenticola (Torrenticola) kurtvietsiMexico stateStreamCramer 1992Neotorrenticola (Neotorrenticola) davecookiMexico stateStreamCramer 1987Pseudotorrenticola (Pseudotorrenticola)VeracruzStreamCramer and Cook 2000Pseudotorrenticola (Psedotorrenticola) boettgeriVeracruzStreamViets 1977 in Cramer and Cook 2000LimnesiidaeNeomamersa (Neomamersa) mexicanaOaxaca, GuerreroBottom deposits of streamCook 1980Neomamersa (Neomamersa) triacebulataChiapasGravels deposits of streamCook 1980Neomamersa (Neomamersa) temazcalaMexico stateStreamCramer 1987Meramecia (Meramecia)mexicanaChiapas, Oaxaca, PueblaGravel deposits of streamCook 1980Kawamuracarus (Kawamuracarus) expansipesOaxaca, ChiapasGravel deposits of streamCook 1980Kawamuracarus (Kawamuracarus) novusPueblaGravel deposits of streamCook 1980Kawamuracarus (Kawamuracarus) iansmithiMexico stateStreamCramer 1987Tyrrelia viudaPueblaStreamCook 1980Tyrrelia ovalisPuebla, Mexico stateStreamMarshall 1940 in Cook 1980Centrolimnesia (Centrolimnesia)motasiChiapasPondCook 1980Centrolimnesia (Centrolimnesia) bondiVeracruz, CampecheSwam, PondLundblad 1935 in Cook 1980Limnesia alzateiGuanajuato*Dugés 1884Limnesia laetaOaxaca, TabascoPondStoll 1887 in Cook 1980Limnesia puteorumGuanajuato*Stoll 1987 in Wolcott 1903Limnesia (Limnesia) abendaMexico stateHelocrene springCook 1980Limnesia (Limnesia) banolaNayarit, VeracruzSpringCook 1980Limnesia (Limnesia) canvadaNayaritRiverCook 1980Limnesia (Limnesia) dimorphaNayaritSpringCook 1980Limnesia (Limnesia) dorneraVeracruzStreamCook 1980Limnesia (Limnesia) mesoamericanaGuerrero, Oaxaca, Chiapas, Morelos, NayaritRiver, streamCook 1980Limnesia (Limnesia) neodentipalpisVeracruzPondCook 1980Limnesia (Limnesia) setacoxalisVeracruz, Chiapas, GuerreroStreamCook 1980Limnesia (Limnesia) trehonaGuerreroSpringCook 1980Limnesia (Limnesia) zomplinaNayarit, VeracruzPondCook 1980Limnesia (Limnesiella) mexicanaTabascoPondCook 1980Limnesia parafalsificataTabasco, ChiapasPondOtero-Colina 1987aPsammolimnesia (Psammolimnesia) mexicanaChiapasCook 1974OmartacaridaeOmartacarus elongatusPueblaInterstitial watersCook 1963 in Cook 1980Omartacarus brevipalpisChiapasInterstitial watersCook 1974Omartacarus motasiChiapasSand and gravel of streamCook 1980HygrobatidaeCorticacarus novumVeracruzStreamCramer and Cook 1998Corticacarus (Paracorticacarus) mexicanusGuerrero, Oaxaca, ChiapasStreamCook 1980Corticacarus (Polycorticacarellus) similisGuerreroStreamCook 1980Corticacarus (Polycorticacarellus) mebosusChiapas, VeracruzRiverCook 1980Neoatractides curvispinatusTabascoGravel in a streamOtero-Colina 1987aNeotractides mexicanaMexico stateStreamCramer 1992Neoatractides nayaritensisNayaritStreamCramer and Cook 2000Neoatractides abueloiVeracruzStreamCramer and Cook 2000Neoatractides celiaeVeracruzStreamCramer and Cook 2000Hygrobates (Hygrobates) boettgeriOaxaca, Chiapas, Guerrero, Morelos, NayaritStreamViets 1975 in Cook 1980Hygrobates (Hygrobates) plebejusOaxaca, Chiapas, Veracruz, Morelos, Nayarit.RiverLundblad 1930 in Cook 1930Hygrobates (Hygrobates) amplipalpisOaxaca, VeracruzStreamViets 1936 in Cook 1980Hygrobates (Hygrobates) ampliatusVeracruz, San Luis Potosí, MorelosStreamViets 1936 in Cook 1980Hygrobates (Hygrobates) procursusGuerreroStreamViets 1936 in Cook 1980Hygrobates (Hygrobates) blatolusOaxacaTributary riverCook 1980Hygrobates (Hygrobates) mexicanusNayaritRiverCook 1980Atractides (Atractides) crassitarsisGuerreroStreamLundblad 1942 in Cook 1980Atractides (Atractides) imitatusOaxacaRiverViets 1978 in Cook 1980Atractides (Atractides) guatemaltecusOaxaca, Chiapas, Guerrero.Tributary river, streamViets 1978 in Cook 1980Atractides (Atractides) blazonusOaxacaTributary riverCook 1980Atractides (Atractides) mexicanusOaxacaTributary riverCook 1980Atractides (Atractides) neolongitarsusCoahuilaRiverCook 1980Atractides (Atractides) prasadiChiapasStreamCook 1980Atractides (Atractides) tanutusMexico stateStreamCook 1980Atractides (Atractides) tembolusChiapasStreamCook 1980Atractides (Atractides) tolasChiapasStreamCook 1980Atractides (Atractides) toldomusCoahuilaRiverCook 1980Atractides (Atractides) travanusChiapas, Oaxaca, GuerreroStreamCook 1980Atractides (Atractides) wevamusNayaritRiver bottom depositsCook 1980Atractides (Atractides) zempoalusMexico stateStreamCook 1980Atractides (Atractides) bassolsaeOaxacaStreamOtero-Colina 1987aAtractides (Octomegapus) peltatusChiapasRiverViets 1977 in Cook 1980Atractidella obtusidensOaxaca, Chiapas, VeracruzRiver, streamLundblad 1953 in Cook 1980Atractidella mesoamericanaChiapasStreamCook 1980Paraschizobates scutatusNayarit, Chiapas, OaxacaRiver, streamCook 1980Diamphidaxona mexicanaChiapasGravel deposits in streamCook 1980Diamphidaxona anitaeMexico stateStreamCramer and Letechipía 1996UnionicolidaeUnionicola gracilipalpis tenuisOaxaca, CampechePond, water filled roadsideLundblad 1935 in Cook 1980Unionicola (Unionicola) mexicanaTabascoPondCook 1980Unionicola (Pentatax) conjunctaChiapasStreamViets 1954 in Cook 1980Unionicola (Pentatax) furculopsisOaxacaPondCook 1980Unionicola (Pentatax) aculeataTamaulipas, northern and central MexicoFreshwater mussel Anodonta sp., Cyrtonaias tampicoensis, Disconaias sp., Friersonia sp.Koenike 1914 in Vidrine 1996Unionicola (Atacella) neoperforataTamaulipas, San Luis Potosí.Freshwater mussel A. trapesialis glaucusVidrine 1985bUnionicola (Atacella) petitaVeracruzFreshwater mussel A. trapesialis glaucusVidrine 1985bUnionicola (Atacella) entrerrianensisTamaulipas, San Luis Potosí, Veracruz.Freshwater mussel A. trapesialis glaucusRosso de Ferradás 1976Unionicola (Atacella) fissipesWestern MexicoFreshwater mussel AnodontitesKoenike, 1891Unionicola (Berezatax) berezaiVeracruz, San Luis PotosíFreshwater mussel Nephronaias sp.Vidrine 1985aUnionicola (Berezatax) acylindrotarsaTamaulipas, San Luis PotosíFreshwater mussel Popenaias, NephronaiasVidrine 1985aUnionicola (Causeyatax) hensleyiSan Luis Potosí, TamaulipasFreshwater mussel Friersonia iridellaVidrine 1985cUnionicola (Clarkatax) serrataSan Luis Potosí, Tamaulipas, Veracruz (Panuco river system)Friersonia iridella, F. moctezumensisWolcott 1898 in Vidrine 1996Unionicola (Hexatax) laurentianaSan Luis Potosí, Tamaulipas, Veracruz (Panuco river system)Freshwater mussel Disconaias fimbriata, Nephronaias sp., Popenaias sp.Crowell and Davids 1979 inUnionicola (Unionicolides) burchiGuanajuatoFreshwater mussel Anodonta sp.Vidrine 1996Unionicola (Unionicolides) calnaniTamaulipas, VeracruzFreshwater mussel Cyrtonaias tampicoensis, Disconaias discus.Vidrine 1986Unionicola (Unionicolides) hoeseiSan Luis Potosí, Tamaulipas, Veracruz (Panuco river system)Freshwater musselVidrine 1986Unionicola (Neoatax) abnormipesTamaulipasFreshwater mussel Cyrtonaias tampicoensisVidrine 1986Neumania (Neumania) conroyiMexico stateReocrene springCook 1980Neumania (Neumania) diversipalpaChiapasRiverCook 1980Neumania (Neumania) alticolaChiapas, Campeche, Oaxaca, Tabasco, Veracruz.Pond, stream.Stoll 1886 in Cook 1980Neumania (Tetraneumania) nondandaChiapas, Tabasco, CampecheRiverCook 1980Neumania (Tetraneumania) cesatecaTabasco, ChiapasRiver, PondOtero-Colina 1987bRecifella (Recifella) bellaVeracruz, ChiapasRiver, streamCook 1980Recifella (Eorecifella?) aztecaOaxacaPondCook 1980Recifella (Eorecifella) cemobaChiapasStreamCook 1980Recifella (Eorecifella) laversiChiapasRiverCook 1980Recifella (Eorecifella) alpaVeracruzStreamCook 1980Recifella (Eorecifella) indistinctaYucatánCenote (Sinkhole)Marshall 1936 in Cook 1980Recifella (Recifellopsis) mexicanaChiapas, VeracruzStreamCook 1980Recifella (Vietsella)veracruzanaVeracruzPondCramer and Cook 1998Koenikea (Koenikea) canermaTabascoPondCook 1980Koenikea (Koenikea) clazonaVeracruzStreamCook 1980Koenikea (Koenikea) gracilipesNayarit, Chiapas, GuerreroStreamCook 1980Koenikea (Koenikea) tobaranaNayaritRiver, streamCook 1980Koenikea (Koenikea) trocalaGuerreroSpringCook 1980Koenikea (Koenikea) yenonaNayaritTributary riverCook 1980Koenikea (Koenikea) zedimaChiapas, TabascoRiverCook 1980Koenikea (Koenikea) zoladaVeracruzRiverCook 1980Koenikea mesoamericanaChiapasStreamCook 1980Koenikea (?Diplokoenikea) bispinaVeracruzRiverCook 1980Koenikea (?Diplokoenikea) bleptaGuerreroStreamCook 1980Koenikea (?Diplokoenikea) beltistaChiapas, Veracruz, Puebla.RiverCook 1980Koenikea (Diplokoenikea) neopectiniferaCampeche, TabascoWaterfilled ditch, pondCook 1980Koenikea (Diplokoenikea) clavigeraChiapas, OaxacaPondLundblad 1943 in Cook 1980Koenikea (Notomideopsis) paragrossaOaxacaPondCook 1980Koenikea (Notomideopsis) sexmaculataOaxaca, VeracruzPondCook 1980Koenikea (Notomideopsis) tolomaVeracruzStreamCook 1980Koenikea (Notomideopsis) veracruzensisVeracruzStreamCook 1980Koenikea (Notomideopsis) nayaritensisNayaritSpringCook 1980Koenikea (Notomideopsis) taninulensisSan Luis PotosíSulphurous streamCramer 1983Koenikea (Sespekoenikea) chiapasanaChiapasStreamCook 1980Koenikea (Sespekoenikea) piotiformisNayarit, GuerreroRiverCook 1980Koenikea (Tanaognathus) kurtvietsiGuerreroRiverCook 1980Koenikea (Parakoenikea) curvipalpisOaxacaRiverLundblad 1936 in Cook 1980PionidaePiona bromadaTabasco, Campeche, Veracruz, Oaxaca.Pond, waterfilled ditchCook 1980Piona mexicanaVeracruzStreamCook 1980Piona neoacutidensVeracruzStreamCook 1980Piona (Piona) triangularis**Wolcott1902 in Marín-Hernández and Cramer-Hemkes 2009Piona (Tetrapiona) rotunda sudamericana**Viets 1910 in Marín-Hernández and Cramer-Hemkes 2009Piona alzatei**Dugés 1884Piona pearsei**Marshall 1936Piona amimitliMexico cityStreamMarín-Hernández 2009AturidaeAturus matlatzincaMexico stateStreamCramer 1991Aturus catoantoniMexico stateStreamCramer 1991Aturus guillecrameriMexico stateStreamCramer 1992Aturus celiahemkesaeMexico stateStreamCramer 1992Aturus oxtotilpanensisMexico stateStreamCramer 1992Aturus primitivusMexico stateStreamCramer 1991Neoaturus kurtvietsiOaxaca, Chiapas, Veracruz, GuerreroRiver, streamCook 1980Neoaturus projectusVeracruzRiverLundblad 1936 in Cook 1980Adelaxonopsella pallidaVeracruzInterstitial water streamCook 1974Albia (Anchistalbia) ianthaNayaritSpringCook 1980Albia (Anchistalbia) ianthopsisQuintana RooLagoonOtero-Colina 1987bAxonopsella (?) mesoamericanaGuerrero, Morelos, NayaritRiverCook 1980Axonopsella (Neoaxonopsella) nayaritensisNayaritCataractsCook 1980Axonopsella (Neoaxonopsella) vicinaVeracruzStreamCook 1980Axonopsis (Brachypodopsis) columbicolaChiapas, Oaxaca, Puebla. NayaritRiver, streamLundblad 1953 in Cook 1980Axonopsis (Brachypodopsis) cemobaVeracruzStreamCook 1980Axonopsis (Brachypodopsis) ceruchaVeracruz, San Luis PotosíRiver, StreamCook 1980Axonopsis (Brachypodopsis) mesoamericanaVeracruz, ChiapasStreamCook 1980Axonopsis (Brachypodopsis) trocalaChiapasChiapasCook 1980Polyaxonopsella mexicanaOaxacaStreamOtero-Colina 1987bFrontipodopsis mesoamericanaChiapasBottom deposits of streamCook 1980Kongsbergia antoniocatoiMexico stateGravel deposits of streamCramer 2000Kongsbergia cositaMexico stateGravel deposits of streamCramer 2000Kongsbergia tigrinaMexico stateStreamCramer 2000Kongsbergia eternaMexico stateStreamCramer 2000Kongsbergia (Kongsbergia) mexicanaOaxacaRiverCook 1980Kongsbergia glopipalpisChiapas and VeracruzStreamLundblad 1953 in Cook 1980Kongsbergia maternaChiapas, Mexico state, Oaxaca, Veracruz.Tributary riverThor 1899 in Cramer 2000Submiraxona (Submiraxona) bellaNayaritStreamCook 1980Submiraxona (Submiraxona) lundbladiGuerrero, ChiapasRiverCook 1980Submiraxona (Submiraxona) stolliGuerreroRiverCook 1980Miraxona (Miraxona) expansipesGuerreroRiverCook 1980Miraxona (Miraxona) mexicanaGuerreroRiverCook 1980Miraxonides (Miraxonidella) similisNayarit, VeracruzRiverCook 1980Miraxona (Miraxonides) geronimoiNayaritStreamCramer and Cook 1991Stygalbiella mexicanaMexico stateSpringCramer 1992Neoaxona (?Lamellaxona) mexicanaNayaritStreamCook 1980Woolastookia gretaeNayaritRiver, streamViets 1978 in Cook 1980Forelia mesoamericanaChiapasLagoonOtero-Colina 1987bFeltriidaeFeltria (Tropifeltria) neotropicaChiapasWaterfallsOtero-Colina 1987bFeltria (Feltriella) anahoffmannaeMexico stateMountain springCramer 1986MideopsidaeMideopsis (Mideopsis) cartesaNayarit, VeracruzStreamCook 1980Mideopsis (Mideopsides) betaChiapasGravel deposits of streamCook 1980Mideopsis (Mideopsis) magnaVeracruz, MorelosRiverCook 1980Mideopsis (Neoxystonotus) nobilisTabascoPondCook 1980Mideopsis orbicularisYucatánCenoteMüller 1776Mideopsis (Neoxystonotus) mexicanaMexico stateHelocrene springCook 1980Mideopsis (Neoxystonotus) gennadaChiapasRiverCook 1980Mideopsis (Neoxystonotus) canzollaGuerreroTributary riverCook 1980KrendowskiidaeKrendowskia (Krendowskiella) aztecaGuerreroTributary riverCook 1980Krendowskia (Krendowskiella) moyaraChiapasRiverCook 1980Krendowskia (Krendowskiella) trodromaVeracruz, NayaritStreamCook 1980Krendowskia (Krendowskiella) vicinaTabascoPondCook 1980Geayia (Geayia) amacuzacaMorelosRiverCook 1980Geayia (Geayia) mitchelliTabasco, ChiapasPondCook 1980ArenohydracaridaeArenohydracarus eremitus**Cook 1974Arenohydracarus minimusOaxaca, GuerreroIntertitial watersCook 1974ChappuisididaeChappuisides notialisMexico stateSpringCramer and Smith 1991HungarohydracaridaeStygarrenurus armoniensisMexico stateInterstitial deposits of spring.Cramer and Cook 1996NeoacaridaeNeoacarus adocetusMexico stateSpringCramer and Smith 1991