five

APPENDIX TABLE 1 in An annotated checklist of the Collembola (Hexapoda) from Iran

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APPENDIX TABLE 1. Taxonomy, Habitat, Ecomorphological life form, Endemic or Exotic, Localities/Provinces and General distribution of Iranian Collembola. Taxonomy/order/ superfamily /subfamily/family/ tribe/genus/speciesHabitatEcomorphological life formEndemic or ExoticLocalities/ProvincesGeneral distributionCLASS COLLEMBOLAORDER PODUROMORPHASUPERFAMILY PODUROIDEAFAMILY PODURIDAEGenus PODURA Linnaeus, 1758Podura aquatica Linnaeus, 1758soil and leaf litterHemiedaphicExoticCentral, TehranCosmopolitanSUPERFAMILY HYPOGASTRUROIDEAFAMILY HYPOGASTRURIDAEGenus ACHERONTIELLA Absolon, 1913Acherontiella bougisi Cassagnau & Delamare Deboutteville, 1955wheat fieldsEpedaphicExoticKhuzestanIndia and IranGenus ACHEROXENYLLA Ellis, 1976Acheroxenylla sp.soilEuedaphic-Kerman-Genus CERATOPHYSELLA Börner, 1932Ceratophysella alani (Babenko 1994)soilEuedaphicExoticGolestanPalaearcticCeratophysella armata (Nicolet, 1842)soilEuedaphicExoticKohgiluyeh and Boyer-Ahmad, Ker- manshahHolarcticCeratophysella cf. borealis Martynova, 1977surface layer of soilEpedaphicExoticLorestanPalaearcticCeratophysella denticulata (Bagnall, 1941)soil and litterHemiedaphicExoticCentral, Mazandaran, Gilan, Ilam, East Azerbaijan, West Azerbaijan, Kerman, Kermanshah, Zanjan, GolestanPalaearctic and NearcticCeratophysella engadinensis (Gisin, 1949)soil and litterHemiedaphicExoticMazandaran. Tehran, Isfahan, IlamCosmopolitanCeratophysella gibbosa (Bagnall, 1940)surface layer of soilEpedaphicExoticKermanshah, Lorestan, North Khorasan, Tehran, KhuzestanCosmopolitanCeratophysella stercoraria Stach, 1963surface layer of soilEpedaphicExoticKermanshah, Kohgiluyeh and Boyer- Ahmad, Mazandaran, Golestan, Tehran, Kerman, LorestanPalaearcticCeratophysella tuberculata Cassagnau, 1959soilEuedaphicExoticKhuzestanPalaearcticGenus CHOREUTINULA Paclt, 1944Choreutinula inermis (Tullberg, 1871)soil and litterHemiedaphicExoticGilanPalaearcticGenus HYPOGASTRURA Bourlet, 1839Hypogastrura assimilis (Krausbauer, 1898)soil and leaf litterHemiedaphicExoticKermanshah, TehranCosmopolitanHypogastrura cf. distincta (Axelson, 1902)soil and leaf litterHemiedaphicExoticTehranHolarctic, AustraliaHypogastrura ellisi Skarżyński, Kahrarian, Piwnik & Zawisza, 2017litterEpedaphicExoticKermanshah, LorestanIran and GreeceHypogastrura manubrialis (Tullberg, 1869)chicken manure, soil and leaf litterHemiedaphicExoticCentral, Kohgiluyeh and Boyer-Ahmad, Mazandaran, Markazi, Gilan, East Azerbaijan, West Azerbaijan, Khuzestan, Zanjan, Tehran, Kermanshah, GolestanCosmopolitanHypogastrura martiani Skarżyński & Kaprus’, 2009soil and litterHemiedaphicExoticKermanshah, MazandaranPalaearcticHypogastrura neglecta (B̂rner, 1901)soilEuedaphicExoticMazandaran,PalaearcticHypogastrura papillata Gisin, 1949leaf litterHemiedaphicExoticMazandaranEurope and IranHypogastrura persica Kahrarian, Vafaei-Shoushtari, Skarżyński & Konikiewicz, 2013soil and litterHemiedaphicEndemicKermanshahIranHypogastrura purpurescens (Lubbock, 1868)soilEuedaphicExoticKermanshah, Tehran, Mazandaran, LorestanCosmopolitanHypogastrura cf. rangkuli Martynova, 1975soil and litterHemiedaphicExoticTehranUncertainHypogastrura serrata (Ågren, 1904)soil and litterHemiedaphicExoticTehranHolarcticHypogastrura socialis (Uzel, 1891)soil and litterHemiedaphicExoticKermanshahPalaearcticHypogastrura cf. tullbergi (Schäffer, 1900)soilEuedaphicExoticMazandaranCircumpolarHypogastrura vernalis (Carl, 1901)soilEuedaphicExoticKohgiluyeh and Boyer-Ahmad, Ma- zandaran, Tehran, North Khorasan, KurdistanCosmopolitanGenus MESACHORUTES Absolon, 1900Mesachorutes sp.soil and litterHemiedaphic-North Khorasan-Genus SCHOETTELLA Schäffer, 1896Schoetella ununguiculata (Tullberg, 1869)soil and mossHemiedaphicExoticMazandaranHolarcticGenus TRIACANTHELLA Schäffer, 1897Triacanthella intermedia Dunger & Zivadinovic, 1984soil and litterHemiedaphicExoticTehran, Mazandaran, North KhorasanBosnia & Herzegovina and IranGenus WILLEMIA Börner, 1901 Willemia anophthalma B̂rner, 1901soilEuedaphicExoticEast Azerbaijan, TehranHolarcticWillemia buddenbrocki Ḩther, 1959soil and litterHemiedaphicExoticKermanshahPalaearcticWillemia denisi Mills, 1932soil and litterHemiedaphicExoticGilanHolarcticWillemia scandinavica Stach, 1949soil and litterHemiedaphicExoticKermanshah, LorestanHolarcticWillemia similis Mills, 1934soil and litterHemiedaphicExoticNorth KhorasanHolarcticWillemia virae Kaprus’, 1997soil and litterHemiedaphicExoticKurdistanUkraine, Poland, Slovak Republik, IranGenus XENYLLA Tullberg, 1869Xenylla humicola (Fabricius, 1780)soil and litterHemiedaphicExoticEast Azerbaijan, West Azerbaijan, GilanHolarcticXenylla maritima Tullberg, 1869mossEpedaphicExoticEast Azerbaijan, West Azerbaijan, Gilan, MazandaranCosmopolitanXenylla mediterranea da Gama, 1964mossEpedaphicExoticMazandaranPalaearcticXenylla mucronata Axelson, 1903soil and litterHemiedaphicExoticNorth KhorasanPalaearcticXenylla szeptyckii Skarżyński, Piwnik & Porco, 2018soil and litterHemiedaphicExoticMazandaranPoland and IranXenylla tullbergi B̂rner, 1903soil and litterHemiedaphicExoticKurdistanPalaearcticXenylla welchi Folsom, 1916soil and litterHemiedaphicExoticMazandaran, Kohgiluyeh and Boyer- Ahmad, IlamPalaearcticSUPERFAMILY ONYCHIUROIDEAFAMILY ONYCHIURIDAESubfamily OnychiurinaeTribe HymenaphoruriniGenus HETERAPHORURA Bagnall, 1948Heteraphorura iranica Kaprus’, Shayanmehr & Kahrarian, 2017soil and litterHemiedaphicEndemicGolestan, Mazandaran, KermanshahIranHeteraphorura cf. japonica (Yosii, 1967)soil and litterHemiedaphicExoticTehran, Mazandaran, Kermanshah, GolestanPalaearcticHeteraphorura kaprusi Arbea, Yahyapour & Shayanmehr, 2020soil and litterHemiedaphicEndemicMazandaranIranGenus HYMENAPHORURA Bagnall, 1948Hymenaphorura prox. sibirica (Tullberg, 1876)soilEuedaphicExoticCentral, Mazandaran, Gilan, Tehran, Zanjan, East Azerbaijan, West Azerbai- janPalaearcticTribe OligaphoruriniGenus OLIGAPHORURA Bagnall, 1949 Oligaphorura absoloni (B̂rner, 1901)soil and litterHemiedaphicExoticKermanshahHolarcticTribe OnychiuriniGenus DEUTERAPHORURA Absolon, 1901Deuteraphorura dashtenazensis Arbea, Yahyapour & Shayanmehr, 2020soil and litterHemiedaphicKnown only from the type localityMazandaranIranGenus ONYCHIUROIDES Bagnall, 1948Onychiuroides cf. granulosus (Stach, 1930)soilEuedaphicExoticCentral, East Azerbaijan, West Azerbai- jan, Gilan, Tehran, Mazandaran, ZanjanPalaearcticOnychiuroides mazandaranensis Arbea, Yahyapour & Shayanmehr, 2020soil and litterHemiedaphicEndemicMazandaranIranOnychiuroides cf. pseudogranulosus (Gisin, 1951)soilEuedaphicExoticCentral, East Azerbaijan, West Azerbai- jan, Gilan, Tehran, Mazandaran, ZanjanPalaearcticGenus ONYCHIURUS Gervais, 1841Onychiurus sp.soilEuedaphic-Mazandaran-Genus ORTHONYCHIURUS Stach, 1954Orthonychiurus folsomi (Schäffer, 1900)soilEuedaphicExoticTehran, Mazandaran, KermanshahPalaearcticOrthonychiurus cf. rectopapillatus (Stach, 1933)soilEuedaphicExoticCentral, East Azerbaijan, West Azerbai- jan, Gilan, Tehran, Mazandaran, ZanjanPalaearcticOrthonychiurus stachianus (Bagnall, 1939)soilEuedaphicExoticTehran, Mazandaran, KermanshahPalaearcticGenus VIBRONYCHIURUS Pomorski, 1998Vibronychiurus archivari (Christiansen, 1956)soil and litterHemiedaphicExoticKermanshahLebanon and IranVibronychiurus hermonicus (Gruia, Poliakov & Broza, 2000)soilEuedaphicExoticKermanshahIsrael and IranTribe ProtaphoruriniGenus PROTAPHORURA Absolon, 1901Protaphorura aurantiaca (Ridley, 1880)soil and litterHemiedaphicExoticSemnanPalaearcticProtaphorura bicampata (Gisin, 1956)soilEuedaphicExoticCentral, Mazandaran, Gilan, Tehran, Zanjan, East Azerbaijan, West Azerbai- janPalaearcticProtaphorura cancellata (Gisin, 1956)soil and litterHemiedaphicExoticKermanshahPalaearcticProtaphorura fimata (Gisin, 1952)soil and manureHemiedaphicExoticCentral, East Azerbaijan, West Az- erbaijan, Gilan, Kermanshah, Isfahan, Tehran, Mazandaran, LorestanPalaearcticProtaphorura golestanica Kaprus’, Shayanmehr & Kahrarian, 2017soil and litterHemiedaphicEndemicGolestan, MazandaranIranProtaphorura gisini (Haybach, 1960)soilEuedaphicExoticLorestanPalaearcticProtaphorura hyrcanica Arbea, Yahyapour & Shayanmehr, 2020soil and litterHemiedaphicKnown only from type localityMazandaranIranProtaphorura iranica Arbea, Yahyapour & Shayanmehr, 2020soil and litterHemiedaphicKnown only from type localityMazandaranIranProtaphorura levantina (Christiansen, 1956)soil and litterHemiedaphicExoticGolestan, Ilam, Kermanshah KurdistanPalaearcticProtaphorura ombrophila (Stach, 1960)soilEuedaphicExoticMazandaranEastern PalaearcticProtaphorura persica Arbea & Kahrarian, 2017soilEuedaphicEndemicKermanshahIranProtaphorura cf. quadriocellata (Gisin, 1947)soil and litterHemiedaphicExoticCentral, East Azerbaijan, West Az- erbaijan, Gilan, Tehran, Mazandaran, Golestan, ZanjanPalaearcticProtaphorura sakatoi (Yosii, 1966)soil and litterHemiedaphicExoticMazandaran, Golestan, KermanshahPalaearcticProtaphorura cf. salsa Kaprus’, Paśnik & Weiner, 2014soil and litterHemiedaphicExoticGolestanRussia and IranTribe ThalassaphoruriniGenus ALLONYCHIURUS Yoshii, 1995Allonychiurus sp.soil and litterHemiedaphic-Golestan-Genus SENSILLONYCHIURUS Pomorski & Shveenkova, 2006Sensillonychiurus sp.soilEuedaphic-Golestan-Genus SPINONYCHIURUS Weiner, 1996Spinonychiurus persicus Kaprus’ Shayanmehr & Kahrarian, 2017soil and litterHemiedaphicEndemicKermanshahIranSpinonychiurus tianshanicus (Martynova, 1971)soilEuedaphicExoticKermanshahKyrgyzstan and IranGenus THALASSAPHORURA Bagnall, 1949Thalassaphorura encarpata (Denis, 1931)soil and litterHemiedaphicExoticMazandaran, Tehran, Lorestan, GolestanCosmopolitanThalassaphorura franzi (Stach, 1946)soil and litterHemiedaphicExoticKurdistanAustria, Italy, Turkey and IranThalassaphorura zschokkei (Handschin, 1919)soil and litterHemiedaphicExoticKermanshah, Lorestan, Ilam, KurdistanNordic speciesFAMILY TULLBERGIIDAEGenus DOUTNACIA Rusek, 1974Doutnacia xerophila Rusek, 1974soil and litterHemiedaphicExoticKurdistanPalaearcticGenus FISSURAPHORURA Rusek, 1991Fissuraphorura duplex (Gama, 1962)soil and litterHemiedaphicExoticKermanshahPalaearcticGenus MESAPHORURA Börner, 1901Mesaphorura critica Ellis, 1976soilEuedaphicExoticGolestanPalaearcticMesaphorura hylophila Rusek, 1982soil and litterHemiedaphicExoticKermanshahPalaearcticMesaphorura italica (Rusek, 1971)soil and litterHemiedaphicExoticKermanshah, KurdistanPalaearcticMesaphorura krausbaueri B̂rner, 1901soilEuedaphicExoticCentral, East Azerbaijan, West Azerbai- jan, Gilan, Tehran, Mazandaran, ZanjanCosmopolitanMesaphorura macrochaeta Rusek, 1976soil and litterHemiedaphicExoticTehran, KermanshahCosmopolitanMesaphorura yosii (Rusek, 1967)soilEuedaphicExoticGolestan, KurdistanCosmopolitanGenus METAPHORURA Stach, 1954 Metaphorura affinis (B̂rner, 1902)soil and litterHemiedaphicExoticCentral, West Azerbaijan, Gilan, Tehran, Kermanshah, IlamPalaearcticMetaphorura denisi Simón, 1985soil and litterHemiedaphicExoticKermanshah, LorestanPalaearcticMetaphorura riozoi Castaño-Meneses, Palacios-Var- gas & Traser, 2000soil and litterHemiedaphicExoticIlam, KurdistanPalaearcticGenus PARATULLBERGIA Womersley, 1930 Paratullbergia callipygos (B̂rner, 1902)soilEuedaphicExoticCentral, Tehran, Zanjan, North Kho- rasanHolarcticParatullbergia macdougalli Bagnall, 1936soil and litterHemiedaphicExoticNorth KhorasanPalaearcticGenus TULLBERGIA Lubbock, 1876Tullbergia simplex Gisin, 1958mossEpedaphicExoticMazandaranHolarcticSUPERFAMILY NEANUROIDEAFAMILY BRACHYSTOMELLIDAE Genus BRACHYSTOMELLA Ågren, 1903Brachystomella nubila Gisin, 1957soilEuedaphicExoticEast Azerbaijan, Gilan, ZanjanGermany and IranBrachystomella parvula (Schäffer, 1896)soil and litterHemiedaphicExoticEast Azerbaijan, Gilan, Zanjan, Tehran, KermanCosmopolitanFAMILY NEANURIDAESubfamily FrieseinaeGenus FRIESEA Dalla Torre, 1895Friesea afurcata Tullberg, 1869moss, humus, soil and leaf litterHemiedaphicExoticGolestanPalaearcticFriesea claviseta Axelson, 1900soil and litterHemiedaphicExoticTehran, MazandaranCosmopolitanFriesea espunaensis Arbea & Jordana, 1993soil and litterHemiedaphicExoticMazandaranSpain and IranFriesea mirabilis (Tullberg, 1871)soil, leaf litter, moss, stock and rotted woodHemiedaphicExoticCentral, Tehran, East Azerbaijan, Gilan, GolestanCosmopolitanSubfamily PseudachorutinaeGenus AETHIOPELLA Handschin, 1942Aethiopella sp.soilEuedaphic-Mazandaran-Genus ANURIDA Laboulbène, 1865Anurida ellipsoides Stach, 1949soilEuedaphicExoticEast Azerbaijan, Gilan, West AzerbaijanPalaearcticAnurida thalassophila (Bagnall, 1939)soilEuedaphicExoticEast Azerbaijan, Gilan, West AzerbaijanPalaearcticGenus ISRAELIMERIA Weiner & Kaprus’, 2005Israelimeria persica Alijani, Shayanmehr, Skarżyński, Piwnik & Smolis, 2015soilEuedaphicKnown only from the type localityGolestanIranGenus MICRANURIDA Börner, 1901Micranurida sensillata (Gisin, 1953)soilEuedaphicExoticEast Azerbaijan, Gilan, West AzerbaijanPalaearcticGenus PSEUDACHORUTES Tullberg, 1871Pseudachorutes corticicolus (Schäffer, 1896)soil and litterHemiedaphicExoticMazandaran, KurdistanHolarcticPseudachorutes dubius Krausbauer, 1898soil, leaf litter and manureHemiedaphicExoticCentral, Tehran, Gilan, MazandaranPalaearcticPseudachorutes hyrcanicus Kaprus’, Shayanmehr & Yahyapour, 2022soil and litterHemiedaphicKnown only from the type localityMazandaranIranPseudachorutes kurdistanicus Kaprus’, Shayanmehr & Ghobari, 2022soil and litterHemiedaphicKnown only from the type localityKurdistanIranPseudachorutes parvulus B̂rner, 1901soil and litterHemiedaphicExoticCentral, Tehran, Gilan, KermanPalaearcticPseudachorutes subcrassus Tullberg, 1871soil and litterHemiedaphicExoticCentral, Mazandaran, Tehran, Gilan, North KhorasanPalaearcticPseudachorutes vitalii Kaprus’ & Weiner, 2009soil and leaf litterHemiedaphicExoticKurdistanUkraine and IranSubfamily MorulininaeGenus MORULINA Börner, 1906 Morulina verrucosa B̂rner, 1903soilEuedaphicExoticGilanPalaearcticSubfamily NeanurinaeGenus BILOBELLA Caroli, 1912Bilobella aurantiaca (Caroli, 1912)soilEuedaphicExoticCentral, Tehran, Gilan, Mazandaran, East AzerbaijanPalaearcticGenus CRYPTONURA Cassagnau, 1979Cryptonura abmalensis Arbea, Yahyapour & Shayanmehr, 2022soil and leaf litterHemiedaphicKnown only from the type localityMazandaranIranCryptonura dohezarensis Arbea, Yahyapour & Shayanmehr, 2022soil and leaf litterHemiedaphicKnown only from the type localityMazandaranIranCryptonura maxima Smolis, Falahati & Skarżyński, 2012soil and leaf litterHemiedaphicEndemicMazandaran, GolestanIranCryptonura persica Smolis, Falahati & Skarżyński, 2012moss, soil and leaf litterHemiedaphicEndemicMazandaran, Golestan, West Azerbaijan, SemnanIranCryptonura sariensis Arbea, Yahyapour & Shayan- mehr, 2022soil and leaf litterHemiedaphicKnown only from the type localityMazandaranIranGenus DEUTONURA Cassagnau, 1979Deutonura breviseta Smolis & Skarżyński, 2020tree holes, leaves and siftingEpedaphicEndemicMazandaran, GilanIranDeutonura decolorata (Gama & Gisin, 1964)soil and leaf litterHemiedaphicExoticMazandaranPalaearcticDeutonura iranica Smolis & Skarżyński, 2020soilEuedaphicEndemicWest AzerbaijanIranDeutonura persica Smolis, Shayanmehr & Yoosefi Lafooraki, 2018moss, leaf litter and dead woodEpedaphicEndemicMazandaranIranDeutonura sengleti Smolis & Skarżyński, 2020tree holes, leaves, and siftingEpedaphicEndemicMazandaran, GilanIranGenus ENDONURA Cassagnau, 1979Endonura agnieskae Smolis & Skarżyński, 2020tree holes, leaves and siftingEpedaphicEndemicMazandaran, Gilan, SemnanIranEndonura annae Smolis & Skarżyński, 2020tree holes, leaves, humus, and sift- ingEpedaphicEndemicMazandaran, GilanIranEndonura ceratolabralis Smolis, Kahrarian, Piwnik & Skarżyński, 2016litterEpedaphicEndemicKermanshah, MazandaranIranEndonura dentifera Smolis, Skarżyński, Pomorski & Kaprus’, 2007soilEuedaphicExoticKermanshahUkraine and IranEndonura dichaeta Smolis, Kahrarian, Piwnik & Skarżyński, 2016litterEpedaphicEndemicKermanshahIranEndonura longirostris Smolis, Shayanmehr, Kuznetsova & Lafooraki, 2017soil and leaf litterHemiedaphicEndemicMazandaran, GolestanIranEndonura paracentaurea Smolis, Shayanmehr, Kuznetsova & Lafooraki, 2017rooting wood, tree hole, humus and large trees in marshEpedaphicEndemicMazandaran, GilanIranEndonura persica Smolis, Kahrarian, Piwnik & Skarżyński, 2016litterEpedaphicEndemicKermanshahIranEndonura schwendingeri Smolis & Skarżyński, 2020tree trunk, leaves and siftingEpedaphicEndemicGilanIranGenus NEANURA MacGillivray, 1893Neanura deharvengi Smolis, Shayanmehr & Yoosefi- Lafooraki, 2018soil and leaf litterHemiedaphicEndemicMazandaran, GilanIranNeanura muscorum (Templeton, 1836)soil and leaf litterHemiedaphicExoticMazandaran, Gilan, Zanjan, North KhorasanCosmopolitanNeanura persica Arbea, Yahyapour & Shayanmehr, 2022soil and leaf litterHemiedaphicEndemicMazandaranIranGenus PARAVIETNURA Smolis & Kuznetsova, 2018Paravietnura rostrata Smolis & Skarżyński, 2020tree hole, leaves and siftingEpedaphicKnown only from the type localityGilanIranGenus PERSANURA Mehrafrooz Mayvan, Shayanmehr, Smolis & Skarżyński, 2015Persanura hyrcanica Mehrafrooz Mayvan, Shayanmehr, Smolis & Skarżyński, 2015soil and leaf litterHemiedaphicKnown only from the type localityMazandaranIranGenus PROTANURA (Börner, 1906)Protanura papillata Cassagnau & Delamare Debout- teville, 1955litterEpedaphicExoticKermanshahLebanon, Israel and IranGenus THAUMANURA Börner, 1932Thaumanura echinata (Kos, 1940)soil and leaf litterHemiedaphicExoticCentralSlovenia and IranGenus VITRONURA Yosii, 1969Vitronura sp.soilEuedaphic-Mazandaran-FAMILY ODONTELLIDAEGenus AXENYLLODES Stach 1949Axenyllodes bayeri Kseneman, 1935soil and leaf litterHemiedaphicExoticGilan, Hamadan, Kermanshah, East Azerbaijan, TehranEurope and IranAxenyllodes cf. caecus Gisin, 1952soil and leaf litterHemiedaphicExoticKurdistanEurope and IranAxenyllodes monoculatus (Jordana & Ardanaz, 1981)litterEpedaphicExoticKermanshahSpain, Qnary Island and IranGenus SUPERODONTELLA Stach, 1949Superodontella lamellifera (Axelson, 1903)soilEuedaphicExoticEast AzerbaijanCosmopolitanORDER ENTOMOBRYOMORPHASUPERFAMILY ENTOMOBRYOIDEAFAMILY ENTOMOBRYIDAESubfamily EntomobryinaeGenus CALX Christiansen, 1958Calx kailashi Mandal, 2018soil and leaf litterHemiedaphicExoticKurdistanIndia and IranGenus COECOBRYA Yosii 1956 sensu Deharveng, 1990Coecobrya tenebricosa (Folsom, 1902)soil and leaf litterHemiedaphicExoticMazandaran, IsfahanCosmopolitanGenus DREPANURA Schött, 1891Drepanura kirgisica Martynova, 1971soilEuedaphicExoticKermanCosmopolitanGenus ENTOMOBRYA Rondani, 1861 Entomobrya atrocincta Sch ̂tt, 1986soil and leaf litterHemiedaphicExoticMazandaran, Kermanshah, Golestan, TehranNearctic and Western EuropeEntomobrya corticalis (Nicolet, 1842)soil and leaf litterHemiedaphicExoticMazandaran, Gilan, Golestan, North KhorasanPalaearcticEntomobrya handschini Stach, 1922soil and leaf litterHemiedaphicExoticKermanshah, Tehran, KurdistanEurope and IranEntomobrya indica (Baijal, 1955)soilEuedaphicExoticKermanTibetan States, India and IranEntomobrya lanuginosa (Nicolet, 1842)soil and leaf litterHemiedaphicExoticCentral, Mazandaran, Tehran, East Azerbaijan, GilanPalaearcticEntomobrya lindbergi Stach, 1960soil and leaf litterHemiedaphicExoticGilan, Mazandaran, Kermanshah, Tehran,South Palaearctic, Afghanistan, Egypt, UAE, Yemen and IranEntomobrya mesopotamica Rusek, 1981soil and leaf litterHemiedaphicExoticMarkazi, KermanshahIraq and IranEntomobrya multifasciata (Tullberg, 1871)soil and leaf litterHemiedaphicExoticMazandaran, Golestan, KurdistanHolarcticEntomobrya nicoleti (Lubbock, 1868)soil and leaf litterHemiedaphicExoticGolestanPalaearcticEntomobrya nigrocincta Denis, 1923soilEuedaphicExoticMazandaran, KermanshahEurope and IranEntomobrya nivalis (Linneaeus, 1758)soil and leaf litterHemiedaphicExoticGolestanNorthern HolarcticEntomobrya numidica Baquero, Hamra-Kroua and Jordana, 2009soil and leaf litterHemiedaphicExoticGolestanAlgeria and IranEntomobrya quinquelineata Bӧrner, 1901soil and leaf litterHemiedaphicExoticMarkaziEurope and IranEntomobrya obscurella Brown, 1926leaf litterEpedaphicExoticMazandaranAsiaEntomobrya schoetti Stach, 1922soilEuedaphicExoticMazandaran, KermanshahEurope and IranEntomobrya unostrigata Stach, 1930soilEuedaphicExoticTehranMediterranean and AsiaGenus MESENTOTOMA Salmon, 1942Mesentotoma dollfussi (Denis, 1924)soilEuedaphicExoticMazandaranEurope, Algeria, Macaronesia and IranMesentotoma subdollfusi Jacquemart, 1974soil and leaf litterHemiedaphicExoticMazandaran, Golestan,golestan GilanEngland, Tunis and IranGenus PRODREPANURA Stach, 1963Prodrepanura sp.soil and leaf litterHemiedaphic-KermanshahGenus SINELLA Brook, 1882Sinella curviseta Brook, 1882soil and leaf litterHemiedaphicExoticMazandaran, Tehran, KermanCosmopolitanSubfamily LepidocyrtinaeGenus LEPIDOCYRTUS Bourlet, 1839Lepidocyrtus bicoloris Mateos, 2012leaf litterHemiedaphicExoticMazandaranSpain and IranLepidocyrtus cyaneus Tullberg, 1871soil and leaf litterHemiedaphicExoticCentral, Mazandaran, East Azerbaijan, Tehran, Zanjan, GilanHolarcticLepidocyrtus lanuginosus (Gmelin, 1790)soil and leaf litterHemiedaphicExoticCentral, Mazandaran, East Azerbaijan, Gilan, Golestan, North KhorasanCommon with a southern distributionLepidocyrtus ruber Sch ̂tt, 1902soilEuedaphicExoticCentral, Mazandaran, East Azerbaijan, Tehran, GilanCosmopolitanGenus PSEUDOSINELLA Schäffer, 1897Pseudosinella alba (Packard, 1873)soilEuedaphicExoticGilan, KurdistanHolarcticPseudosinella baghdadica Rusek, 1981soilEuedaphicExoticKhuzestan, KermanshahIraq and IranPseudosinella duodecimpunctata Denis, 1931soil and leaf litterHemiedaphicExoticEast AzerbaijanEuropePseudosinella imparipunctata Gisin, 1953soilEuedaphicExoticMazandaran, Gilan, ZanjanHolarcticPseudosinella octopunctata B̂ner, 1901moss, soil and leaf litterHemiedaphicExoticCentral, Zanjan, Gilan, Ilam, East Azerbaijan, Kohgiluyeh and Boyer-Ah- mad, North Khorasan, West Azerbaijan, Markazi, Mazandaran, Tehran, Isfahan, Golestan, Kurdistan, Kermanshah, Lorestan, KermanHolarcticPseudosinella sexoculata Sch ̂tt, 1902soilEuedaphicExoticKermanshah, KurdistanCosmopolitanSubfamily SeirinaeGenus SEIRA Lubbock, 1870Seira atlantica Negri, Pellecchia & Fanciulli, 2005soilEuedaphicExoticZanjan, North KhorasanMorocco and IranSeira domestica (Nicolet, 1842)trunk of the tree, soil and leaf litterHemiedaphicExoticMazandaran, Gilan, Golestan, Kerman- shahEuro-Mediterranean region and IranSeira dori Gruia, Poliakov & Broza, 2000soilEuedaphicExoticKurdistanEuro-Mediterranean region and IranSubfamily WillowsiinaeGenus DREPANOSIRA Bonet, 1942Drepanosira hussi Neuherz, 1976soilEuedaphicExoticNorth KhorasanIndia and IranGenus WILLOWSIA Shoebotham, 1917Willowsia bartkei Stach, 1965soilEuedaphicExoticNorth KhorasanPalaearcticWillowsia nigromaculata (Lubbock, 1873)soilEuedaphicExoticGilanCosmopolitanFAMILY ORCHESELLIDAESubfamily HeteromurinaeGenus HETEROMURTRELLA Mari Mutt, 1979Heteromurtrella sp.soilEuedaphic-North Khorasan-Genus HETEROMURUS Wankel, 1860Heteromurus gigans Mari Mutt & Stomp, 1980leaf litterEpedaphicExoticMazandaran, KurdistanEuropeHeteromurus variabilis (Martynova, 1974)leaf litterEpedaphicExoticMazandaran, North KhorasanTajikistan and IranHeteromurus major (Moniez, 1889)compost and soilHemiedaphicExoticCentral, Mazandaran, Gilan, Tehran, Kermanshah, Golestan, Lorestan, KurdistanEuro-MediterraneanHeteromurus nitidus (Templeton, 1836)soil and leaf litterHemiedaphicExoticMazandaran, North Khorasan, Gilan, Golestan, Kermanshah, Kohgiluyeh and Boyer-AhmadHolarcticHeteromurus sexoculatus Brown, 1926soil and leaf litterHemiedaphicExoticMazandaran, KermanshahAsia MinorSubfamily OrchesellinaeGenus ORCHESELLA Templeton, 1836Orchesella cincta (Linnaeus, 1758)moss, natural compost, soil and leaf litterHemiedaphicExoticMazandaran, North Khorasan, GolestanHolarcticOrchesella flavescens (Bourlet, 1839)soilEuedaphicExoticMazandaranHolarcticOrchesella spectabilis Tullberg, 1871leaf litterEpedaphicExoticMazandaranPalaearcticFAMILY PARONELLIDAE Börner, 1906Subfamily Cyphoderinae Börner, 1906Genus CYPHODERUS Nicolet, 1842 Cyphoderus agnotus B̂ner, 1906soilEuedaphicExoticGolestanBoreal and NeotropicalCyphoderus albinus Nicolet, 1842soil and leaf litterHemiedaphicExoticGilan, Tehran, Khuzestan, Kohgiluyeh and Boyer-Ahmad, Isfahan, Kerman, Mazandaran, Kermanshah, KurdistanPalaearcticCyphoderus bidenticulatus Parona, 1888soil and leaf litterHemiedaphicExoticKermanshah, LorestanPalaearcticSUPERFAMILY ISOTOMOIDEAFAMILY ISOTOMIDAESubfamily AnurophorinaeGenus ANUROPHORUS Nicolet, 1842Anurophorus coiffaiti Cassagnau & Delamare, 1955soilEuedaphicExoticGolestan, Gilan, LorestanPalaearcticAnurophorus silvaticus Potapov & Stebaeva, 1990moss on tree, decaying wood and soilHemiedaphicExoticMazandaranRussia, Caucasus, Azerbaijan and IranGenus FOLSOMIA Willem, 1902Folsomia asiatica Martynova, 1971leaf litterEpedaphicExoticKermanshah, Golestan, LorestanMiddle AsiaFolsomia binoculata (Wahlgren 1899)soil and leaf litterHemiedaphicExoticKermanshah, IlamPalaearcticFolsomia brevifurca (Bagnall, 1949)soil and leaf litterHemiedaphicExoticMazandaranPalaearcticFolsomia candida (Willem, 1902)soil and leaf litterHemiedaphicExoticCentral, Mazandaran, Gilan, Tehran, East Azerbaijan, Golestan, KurdistanCosmopolitanFolsomia diplophthalma (Axelson, 1902)soilEuedaphicExoticKermanPalaearcticFolsomia fimetaria (Linnaeus, 1758)soil and leaf litterHemiedaphicExoticCentral, Mazandaran, Gilan, TehranHolarcticFolsomia inoculata Stach, 1947soil and leaf litterHemiedaphicExoticMazandaranHolarcticFolsomia ksenemani Stach, 1947soil and leaf litterHemiedaphicExoticMazandaran, TehranPalaearcticFolsomia manolachei Bagnall, 1939soilEuedaphicExoticKermanshah, Golestan, KurdistanPalaearcticFolsomia penicula Bagnall, 1939soil and leaf litterHemiedaphicExoticCentral, Lorestan, East Azerbaijan, Go- lestan, Gilan, Tehran, Mazandaran, Ker- manshah, North Khorasan, KurdistanPalaearcticFolsomia quadrioculata (Tullberg, 1871)soil and leaf litterHemiedaphicExoticCentral, Lorestan, East Azerbaijan, Golestan, Gilan, Tehran, Mazandaran, Kermanshah, North Khorasan, Gilan, Ilam, West Azerbaijan, KurdistanHolarcticFolsomia similis Bagnall, 1939soil and leaf litterHemiedaphicExoticGilan, Tehran, KermanHolarcticFolsomia trisetata Jordana & Ardanaz, 1981soil and leaf litterHemiedaphicExoticMazandaranSpain and IranGenus GNATHOFOLSOMIA Deharveng & Christian, 1984Gnathofolsomia sp.soil and leaf litterHemiedaphic-Kermanshah-Genus HEMISOTOMA Bagnall, 1949Hemisotoma orientalis (Stach, 1947)soil and leaf litterHemiedaphicExoticCentral, Mazandaran, Gilan, Tehran, East Azerbaijan, West Azerbaijan, Ker- manshahEurope and IranHemisotoma pontica (Stach, 1947)soilEuedaphicExoticCentral, Mazandaran, Gilan, Tehran, North Khorasan, East Azerbaijan, Go- lestan, West Azerbaijan, Ilam, Kerman- shah, KurdistanCosmopolitanHemisotoma quadrioculata (Martynova, 1968)soilEuedaphicExoticIlam, KurdistanMiddle AsiaHemisotoma thermophila (Axelson, 1900)rotted cow ma- nure, soil and leaf litterHemiedaphicExoticCentral, Mazandaran, Gilan, Tehran, East Azerbaijan, West Azerbaijan, Ker- manshah, GolestanCosmopolitanGenus ISOTOMIELLA Bagnall, 1939Isotomiella gracilimucronata Rusek, 1981soilEuedaphicExoticNorth KhorasanIran and IraqIsotomiella minor (Schäffer, 1896)soil and leaf litterHemiedaphicExoticMazandaran, Gilan, Tehran, East Az- erbaijan, North Khorasan, Kermanshah, Golestan, Lorestan, Kerman, KurdistanHolarcticGenus ISOTOMODES Axelson, 1907Isotomodes golestani Yoosefi Lafooraki, Hajizadeh, Shayanmehr & Hosseini, 2020soilEuedaphicKnown only from the type localityMazandaranIranIsotomodes korkorensis Arbea & Kahrarian, 2015soil and leaf litterHemiedaphicKnown only from the type localityKermanshahIranSubfamily IsotominaeGenus AGRENIA Börner, 1906Agrenia sp.soilEuedaphic-North Khorasan-Genus DESORIA Agassiz & Nicolet in Desor, 1841Desoria neglecta (Schäffer, 1900)soil and leaf litterHemiedaphicExoticLorestan, KurdistanHolarcticDesoria olivacea (Tullberg, 1871)soilEuedaphicExoticEast AzerbaijanHolarcticDesoria tigrina Nicolet, 1842soilEuedaphicExoticKermanshah, LorestanPalaearcticDesoria trispinata (MacGillivray, 1896)soilEuedaphicExoticNorth KhorasanPalaearcticDesoria zlotini Martynova, 1962soil and leaf litterHemiedaphicExoticKermanshahAsiaGenus ISOTOMA Bourlet, 1839Isotoma anglicana Lubbock, 1862soil and leaf litterHemiedaphicExoticIlamHolarcticIsotoma iranica Arbea & Kahrarian, 2015soilEuedaphicKnown only from the type localityKermanshah, GolestanIranIsotoma cf. riparia (Nicolet, 1842)soilEuedaphicExoticMazandaranPalaearcticIsotoma viridis Bourlet, 1839soil and leaf litterHemiedaphicExoticCentral, East Azerbaijan, West Azerbai- jan, Tehran, Mazandaran, Kermanshah, Golestan, KurdistanHolarcticGenus ISOTOMURUS Börner, 1903Isotomurus afghanicus Yosii, 1966soil and leaf litterHemiedaphicExoticMazandaranAfghanistan and IranIsotomurus antennalis (Bagnall, 1940)soilEuedaphicExoticMazandaranPalaearcticIsotomurus cf. balteatus (Reuter, 1876)leaf litterEpedaphicExoticGolestanUnclearIsotomurus cassagnaui Deharveg & Lek, 1993soilEuedaphicExoticMazandaranFrance and IranIsotomurus fucicolus (Scĥtt, 1893)soilEuedaphicExoticTehran, Mazandaran, Gilan, GolestanHolarcticIsotomurus graminis Fjellbeg, 2007soilEuedaphicExoticNorth KhorasanPalaearcticIsotomurus hyrcanicus Yoosefi Lafooraki & Shayanmehr, 2023soil, leaf litter, dead wood and mossHemiedaphicKnown only from the type localityMazandaran, GolestanIranIsotomurus katule Yoosefi Lafooraki & Shayanmehr, 2023soil and leaf litterHemiedaphicKnown only from the type localityGolestanIranIsotomurus maculatus (Schäffer, 1896)soil and leaf litterHemiedaphicExoticGolestan, MazandaranUnclearIsotomurus palustris (M̧ller, 1776)soilEuedaphicExoticCentral, Golestan, Mazandaran, Gilan, Khuzestan, IsfahanUnclearIsotomurus potapovi Yoosefi Lafooraki & Shayan- mehr, 2023soil, leaf litter and mossHemiedaphicKnown only from the type localityMazandaran, Golestan, GilanIranIsotomurus punctiferus Yosii, 1963soil and leaf litterHemiedaphicExoticTehran, Gilan, GolestanAsiaIsotomurus stuxbergi (Tullberg, 1876)soilEuedaphicExoticMazandaranHolarcticGenus PARISOTOMA Bagnall, 1940Parisotoma notabilis (Schäffer, 1896)soil and leaf litterHemiedaphicExoticCentral, East Azerbaijan, West Azerbai- jan, Zanjan, North Khorasan, Tehran, Mazandaran, Kermanshah, Golestan, Gilan, Lorestan, Khuzestan, Kerman, KurdistanCosmopolitanGenus PSEUDISOTOMA Handschin, 1924Pseudisotoma sensibilis (Tullberg, 1876)under dead woodEuedaphicExoticMazandaranPalaearcticGenus VERTAGOPUS Börner, 1906Vertagopus persicus Potapov, Yoosefi & Shayanmehr, 2020mossEpedaphicEndemicMazandaran, GolestanIranSubfamily PachyotominaeGenus PACHYOTOMA Bagnall, 1949Pachyotoma sp.soilEuedaphic-Kermanshah-Subfamily ProisotominaeGenus BAGNALLELLA Salmon, 1951Bagnallella tenella (Reuter, 1895)soilEuedaphicExoticGilan, LorestanEurope and IranGenus BALLISTURA Börner, 1906Ballistura schoetti (Dalla Torre, 1895)soil and leaf litterHemiedaphicExoticGilanCosmopolitanBallistura tuberculata (Stach, 1947)soilEuedaphicExoticKhuzestanHolarcticGenus EPHEMEROTOMA Potapov et al., 2015Ephemerotoma skarzynskii Potapov, Kahrarian, Deharveng & Shayanmehr, 2015soilEuedaphicKnown only from the type localityKermanshahIranGenus FOLSOMIDES Stach, 1922Folsomides angularis (Axelson, 1905)soilEuedaphicExoticGilan, KurdistanHolarcticFolsomides halshinicus Arbea & Kahrarian, 2015soil and leaf litterHemiedaphicEndemicKermanshah, KurdistanIranFolsomides marchicus (Frenzel, 1941)soil and leaf litterHemiedaphicExoticKermanshah, Lorestan, Ilam, KurdistanPalaearcticFolsomides parvulus Stach, 1922soil and leaf litterHemiedaphicExoticCentral, East Azerbaijan, West Az- erbaijan, North Khorasan, Tehran, Mazandaran, Gilan, Semnan, Golestan, Lorestan, Kerman, KurdistanCosmopolitanFolsomides subvinosus Arbea & Kahrarian, 2015soil and leaf litterHemiedaphicEndemicKermanshahIranGenus ISOTOMODELLA Martynova, 1968Isotomodella alticola (Bagnall, 1949)soilEuedaphicExoticLorestanPalaearcticGenus PROISOTOMA Börner, 1903Proisotoma minima Absolon, 1901leaf litterEpedaphicExoticGolestan, MazandaranPalaearcticProisotoma minuta (Tullberg, 1871)soil and leaf litterHemiedaphicExoticCentral, Zanjan, East Azerbaijan, Gilan,West Azerbaijan, Tehran, Mazandaran, Isfahan, GolestanCosmopolitanProisotoma subminuta Denis, 1931soilEuedaphicExoticTehran, Mazandaran, Golestan, GilanCosmopolitanSUPERFAMILY TOMOCEROIDEAFAMILY ONCOPODURIDAEGenus ONCOPODURA Carl & Lebedinsky, 1905Oncopodura ambigua Christiansen, 1957soilEuedaphicExoticMazandaran, GilanLebanon and IranOncopodura hamata Carl & Lebedinsky 1905soilEuedaphicExoticGilanEuropeFAMILY TOMOCERIDAEGenus POGONOGNATHELLUS Paclt, 1944Pogonognathellus flavescens (Tullberg, 1871)soil and leaf litterHemiedaphicExoticMazandaranPalaearctic and NearcticGenus TOMOCERINA Yosii, 1955Tomocerina minuta (Tullberg, 1876)soil and leaf litterHemiedaphicExoticMazandaranHolarcticGenus TOMOCERUS Nicolet, 1842Tomocerus minor (Lubbock, 1862)soil and leaf litterHemiedaphicExoticCentral, Zanjan, East Azerbaijan, Gilan, Golestan, Kerman, MazandaranHolarcticTomocerus vulgaris (Tullberg, 1871)soil and leaf litterHemiedaphicExoticCentral, Zanjan, East Azerbaijan, Gilan, North Khorasan, Tehran, Mazandaran, Golestan, KermanshahHolarcticORDER NEELIPLEONAFAMILY NEELIDAEGenus MEGALOTHORAX Willem, 1900 Megalothorax incertus B̂rner, 1903soil and leaf litterHemiedaphicExoticMazandaran, KermanshahCosmopolitanMegalothorax minimus Willem, 1900soilEuedaphicExoticNorth KhorasanCosmopolitanMegalothorax perspicillum Schneider & D’Haese, 2013dead timbers, soil and leaf litterHemiedaphicExoticMazandaran, KermanshahCosmopolitanMegalothorax willemi Schneider & D’Haese, 2013soilEuedaphicExoticMazandaran, KermanshahBelguim and IranGenus NEELUS Folsom, 1896Neelus murinus Folsom, 1896soil and leaf litterHemiedaphicExoticMazandaran, East Azerbaijan, Tehran, IlamCosmopolitanORDER SYMPHYPLEONASUPERFAMILY DICYRTOMOIDEAFAMILY DICYRTOMIDAESubfamily DicyrtominaeGenus DICYRTOMA Bourlet, 1842Dicyrtoma fusca (Lubbock, 1873)leaf litterEpedaphicExoticMazandaranHolarcticDicyrtoma ghilarovi Bretfeld, 1996soil and leaf litterHemiedaphicExoticMazandaranPalaearcticDicyrtoma grinbergsi Stebaeva, 1966soil and leaf litterHemiedaphicExoticLorestanRussia and TaiwanGenus DICYRTOMINA Börner, 1903Dicyrtomina minuta (O. Fabricius, 1783)soil and leaf litterHemiedaphicExoticMazandaran, GilanHolarcticDicyrtomina ornata (Nicolet, 1842)leaf litterEpedaphicExoticMazandaran, Kohgiluyeh and Boyer- AhmadPalaearcticSUPERFAMILY KATIANNOIDEAFAMILY ARRHOPALITIDAEGenus ARRHOPALITES Börner, 1906Arrhopalites caecus (Tullberg, 1871)soil and leaf litterHemiedaphicExoticGilan, East Azerbaijan, North Khorasan, Zanjan, Mazandaran, KermanshahPalaearcticArrhopalites persicus Vargovitsh & Kahrarian, 2020surface layer of soilEpedaphicEndemicKermanshahIranGenus PYGMARRHOPALITES Vargovitsh, 2009Pygmarrhopalites principalis (Stach, 1945)soil and leaf litterHemiedaphicExoticMazandaranHolarcticFAMILY KATIANNIDAEGenus SMINTHURINUS Börner, 1901Sminthurinus aureus (Lubbock, 1862)soil and leaf litterHemiedaphicExoticNorth Khorasan, Mazandaran, Gilan, KurdistanPalaearcticSminthurinus bimaculatus Axelson, 1902soil and leaf litterHemiedaphicExoticGilanPalaearcticSminthurinus elegans (Fitch, 1863)moss on timbers, soil and leaf litterHemiedaphicExoticNorth Khorasan, Mazandaran, Koh- giluyeh and Boyer-Ahmad, Golestan, Kerman, Kermanshah, Tehran, LorestanHolarcticSminthurinus gisini da Gama, 1956moss on the trunkEpedaphicExoticMazandaranEurope and IranSminthurinus reticulatus Cassagnau, 1964moss and soilHemiedaphicExoticKohgiluyeh and Boyer-AhmadPalaearcticSminthurinus signatus (Krausbauer, 1898)soilEuedaphicExoticKohgiluyeh and Boyer-AhmadPalaearcticSminthurinus transversalis Axelson, 1905moss and soilHemiedaphicExoticKohgiluyeh and Boyer-AhmadPalaearcticSUPERFAMILY SMINTHURIDOIDEAFAMILY SMINTHURIDIDAEGenus SMINTHURIDES Börner, 1900Sminthurides aquaticus (Bourlet, 1842)soilEuedaphicExoticKohgiluyeh and Boyer-AhmadHolarcticSminthurides inaequalis B̂rner, 1903soilEuedaphicExoticMazandaranItaly, Palermo, Portugal and IranSminthurides malmgreni (Tullberg, 1876)soil and leaf litterHemiedaphicExoticCentral, East Azerbaijan, Gilan, TehranHolarcticGenus SPHAERIDIA Linnaniemi, 1912Sphaeridia pumilis (Krausbauer, 1898)soil and leaf litterHemiedaphicExoticCentral, North Khorasan, Gilan, Tehran, Markazi, Lorestan, Mazandaran, Go- lestan, KermanshahHolarcticGenus STENACIDIA Börner, 1906Stenacidia violacea (Reuter, 1881)soilEuedaphicExoticGilanHolarcticSUPERFAMILY SMINTHUROIDEAFAMILY BOURLETIELLIDAEGenus BOURLETIELLA Banks, 1899Bourletiella sp.soilEuedaphic-North Khorasan-Genus HETEROSMINTHURUS Stach, 1955Heterosminthurus insignis (Reuter, 1876)soil and leaf litterHemiedaphicExoticKurdistanPalaearcticFAMILY SMINTHURIDAESubfamily SminthurinaeGenus ALLACMA Börner, 1906Allacma fusca (Linnaeus, 1758)pitfall trapsEpedaphicExoticMazandaranHolarcticGenus CAPRAINEA Dallai, 1970Caprainea marginata (Schott, 1893)soilEuedaphicExoticMazandaran, Kermanshah, North Kho- rasanPalaearcticGenus SMINTHURUS Latreille, 1804Sminthurus ghilarovi Stebaeva, 1966soil and leaf litterHemiedaphicExoticMazandaranPalaearcticSminthurus muscicolus Betsch, 1977soil and leaf litterHemiedaphicExoticIlamRussia, Mongolia and IranSminthurus nigromaculatus Tullberg, 1871soilEuedaphicExoticMazandaran, KermanshahHolarcticSminthurus viridis Linnaeus, 1758soilEuedaphicExoticKhuzestan, Mazandaran, Kermanshah, North Khorasan, LorestanCosmopolitanSminthurus wahlgreni Stach, 1920soilEuedaphicExoticKhuzestanEastern and central Europe and IranSubfamily SphyrothecinaeGenus LIPOTHRIX Börner, 1906Lipothrix lubbocki (Tullberg, 1872)soil and leaf litterHemiedaphicExoticMazandaranPalaearcticGenus NEOSMINTHURUS Mills, 1934Neosminthurus sp.soil and leaf litterHemiedaphic-Golestan-Genus PARALIPOTHRIX Bretfeld, 1999Paralipothrix natalicius (Ellis, 1974)soil and leaf litterHemiedaphicExoticMazandaranPalaearcticGenus SPHYROTHECA Bӧrner, 1906Sphyrotheca sp.soil and leaf litterHemiedaphic-Mazandaran- ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page ......continued on the next page
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