Appendix 1 in Integrative taxonomy supports the description of two new species of Macrobiotidae (Tardigrada, Eutardigrada) from Kristianstads Vattenrike Biosphere Reserve in Sweden

Appendix 1 (continued on next 14 pages). GenBank Accession numbers for COI, 18S, ITS, and 28S sequences of all specimens analysed in this study. New sequences are highlighted in red. TaxonSpecimenCOI18SITS28STaxon name in GenBankNotesReference in GenBankDiaforobiotus islandicusIS.042MT808072MT812470MT812597MT812461Diaforobiotus islandicusStec et al. 2020dMacrobiotus aff. polonicusS165.1MW593929MW588026MW588021MW588032Macrobiotus aff. polonicus S165Vecchi & Stec 2021Macrobiotus aff. polonicusS165.2MW593930MW588027MW588020MW588033Macrobiotus aff. polonicus S165Vecchi & Stec 2021Macrobiotus almadaiZYL123MW990245MW995180Macrobiotus almadaiWang et al. (unpubl.)Macrobiotus annewintersaeS207.1MW593927MW588025MW588019MW588030Macrobiotus annewintersaeVecchi & Stec 2021Macrobiotus annewintersaeS207.2MW593928MW588024MW588018MW588031Macrobiotus annewintersaeVecchi & Stec 2021Macrobiotus ariekammensis ariekammensisNO.393.01MZ460999MZ463668MZ463657MZ463674Macrobiotus ariekammensis ariekammensisStec et al. 2022Macrobiotus ariekammensis groenlandicusGL.018.04MZ461006MZ463664MZ463654MZ463679Macrobiotus ariekammensis groenlandicusStec et al. 2022Macrobiotus azzunaeC4218_T5MW698698Macrobiotus sp. 1 MC-2021temporary taxon name has not been updated in GenBank at time of writingMarnissi et al. 2021Macrobiotus azzunaeC4218_V4MW698697MW695447MW695454MW695450Macrobiotus sp. 1 MC-2021temporary taxon name has not been updated in GenBank at time of writingMarnissi et al. 2021Macrobiotus basiatusUSA /NEL/1MT502116MT498094MT505165MT488397Macrobiotus sp. 1 DG-2020temporary taxon name has not been updated in GenBank at time of writingNelson et al. 2020Macrobiotus birendraiCN8.101MW656266MW680641MW680418MW680644Macrobiotus birendraiKayastha et al. 2021Macrobiotus caelestisKG.007MK737922MK737073MK737072MK737071Macrobiotus caelestisCoughlan et al. 2019Macrobiotus canaricusES.004_H1MH057765MH063925MH063928MH063934Macrobiotus canaricusStec et al. 2018cMacrobiotus canaricusES.004_H2MH057766Macrobiotus canaricusStec et al. 2018cMacrobiotus cf. hufelandiC2959aHQ876590OP596303Macrobiotus cf. hufelandi 1 MC-2011Bertolani et al. 2011Macrobiotus cf. muralisC3251_2OP561785OP596299Macrobiotus cf. muralisBertolani et al. 2023Macrobiotus cf. muralisC3251_3OP561786OP596300Macrobiotus cf. muralisBertolani et al. 2023Macrobiotus cf. muralisC3251_4OP561788Macrobiotus cf. muralisBertolani et al. 2023Macrobiotus cf. muralisC3251_5OP561789Macrobiotus cf. muralisBertolani et al. 2023Macrobiotus cf. muralisC3251_FAOP561787OP596301Macrobiotus cf. muralisBertolani et al. 2023Macrobiotus cf. nelsonae1HQ604965MacrobiotusBertolani et al. 2014Macrobiotus cf. recensES.006 haplotype 1MH057768MH063927MH063932MH063936Macrobiotus cf. recens DS-2018Stec et al. 2018cMacrobiotus cf. sapiensS12.Mac.1OK662997OK663226OK663215OK663237Macrobiotus cf. sapiensVecchi et al. 2022cMacrobiotus cf. shonaicusMM1-2018LC431582LC431591Macrobiotus cf. shonaicusMatsumoto & Sugiura (unpubl.)Macrobiotus crustulusGF.271MT260371MT261912MT261907MT261903Macrobiotus crustulusStec et al. 2020aMacrobiotus dolosusC3209_2OP561772OP596290Macrobiotus dolosusBertolani et al. 2023Macrobiotus dolosusC3209_4OP561774Macrobiotus dolosusBertolani et al. 2023Macrobiotus dolosusC3209_ US 2OP561773Macrobiotus dolosusBertolani et al. 2023Macrobiotus dolosusC3581_V6OP561775OP596292Macrobiotus dolosusBertolani et al. 2023Macrobiotus engbergihaplotype 1MN444824MN443039MN443036MN443034Macrobiotus engbergiVecchi et al. 2024bMacrobiotus engbergihaplotype 2MN444825MN443037Macrobiotus engbergiVecchi et al. 2024bMacrobiotus fontouraiC2861_5OP561783OP596295Macrobiotus fontouraiBertolani et al. 2023Macrobiotus fontouraiC2861_ US 1OP561784OP596296Macrobiotus fontouraiBertolani et al. 2023Macrobiotus glebkaiUA.003MW246134MW247177MW247180MW247176Macrobiotus glebkaiKiosya et al. 2021Macrobiotus hannaePL.010MH057764MH063922MH063923MH063924Macrobiotus hannaeNowak & Stec 2018Macrobiotus hufelandiC2953aHQ876585OP596302Macrobiotus hufelandiBertolani et al. 2023Macrobiotus hufelandiS605.Mac.1OK662992OK663221OK663210OK663232Macrobiotus hufelandiBertolani et al. 2023Macrobiotus joannae1HQ604974Macrobiotus joannaeBertolani et al. 2014Macrobiotus kamilaeIN.030 haplotype 1MK737920MK737070MK737067MK737064Macrobiotus kamilaeCoughlan & Stec 2019Macrobiotus kathyaeUS.006.01PP386934PP391285PP391289PP391287Macrobiotus sp. MV2024temporary taxon name has not been updated in GenBank at time of writingMassa & Vecchi 2024Macrobiotus kathyaeUS.006.02PP386935PP391286PP391290PP391288Macrobiotus sp. MV2024temporary taxon name has not been updated in GenBank at time of writingMassa & Vecchi 2024Macrobiotus kathyaeUS.006.03PP386932Macrobiotus sp. MV2024temporary taxon name has not been updated in GenBank at time of writingMassa & Vecchi 2024Macrobiotus kathyaeUS.006.04PP386933Macrobiotus sp. MV2024temporary taxon name has not been updated in GenBank at time of writingMassa & Vecchi 2024Macrobiotus kirghizicusKG.062.01MZ461002MZ463666MZ463659MZ463672Macrobiotus kirghizicusStec et al. 2022Macrobiotus kosmaliM8.1OP141639OP142472OP153786OP143765Macrobiotus kosmaliKayastha et al. 2023aMacrobiotus kristenseniC3291_A02_V2KC193573KC193577Macrobiotus kristenseniGuidetti et al. 2013Macrobiotus kyoukenusC4313_ US 3ON809462ON818314ON818301ON818309Macrobiotus kyoukenusCesari et al. 2022Macrobiotus macrocalixPL.110MH057767MH063926MH063931MH063935Macrobiotus macrocalixStec et al. 2018cMacrobiotus margoaeUS.057MN888315MN888368MN888340MN888354Macrobiotus margoaeVecchi & Stec 2021Macrobiotus margoaeUS.057.1MT807927MT809072MT809098MT809084Macrobiotus margoaeStec et al. 2021bMacrobiotus margoaeUS.057.3MT807928MT809099Macrobiotus margoaeStec et al. 2021bMacrobiotus mileriIL.001.01OR544397OR543312OR543316OR543320Macrobiotus mileriStec 2024Macrobiotus mileriIL.001.02OR544398OR543313OR543317OR543321Macrobiotus mileriStec 2024Macrobiotus mileriIL.001.03OR544399Macrobiotus mileriStec 2024Macrobiotus mileriIL.001.04OR544400Macrobiotus mileriStec 2024Macrobiotus noongarisAU.031MK737919MK737069MK737066MK737063Macrobiotus noongarisCoughlan & Stec 2019Macrobiotus ovovittatusGL.001.01OR544395OR543311OR543314OR543318Macrobiotus ovovittatusStec 2024Macrobiotus pallariiIT.337.1MT807924MT809069MT809094MT809081Macrobiotus pallariiStec et al. 2021bMacrobiotus pallariiIT.337.2MT807925MT809070MT809095MT809082Macrobiotus pallariiStec et al. 2021bMacrobiotus pallariiIT.337.3MT807926MT809071MT809096MT809083Macrobiotus pallariiStec et al. 2021bMacrobiotus papeiTZ.027MH057763MH063881MH063921MH063880Macrobiotus papeiStec et al. 2018bMacrobiotus paulinaeKT 951668KT935502KT935500Macrobiotus paulinaeStec et al. 2015Macrobiotus scoticusDK.056.1OK662989OK663217OK663206OK663228Macrobiotus scoticusVecchi et al. 2022cMacrobiotus shonaicushaplotype 1MG757136MG757132MG757134MG757133Macrobiotus shonaicusStec et al. 2018aMacrobiotus siderophilusC2796_2OP561776OP596293Macrobiotus siderophilusBertolani et al. 2023Macrobiotus siderophilusC3282_UFKOP561777Macrobiotus siderophilusBertolani et al. 2023Macrobiotus siderophilusC3282_UFLOP561778Macrobiotus siderophilusBertolani et al. 2023Macrobiotus siderophilusC3282_UFMOP561779Macrobiotus siderophilusBertolani et al. 2023Macrobiotus siderophilusC3282_UFNOP561780OP596294Macrobiotus siderophilusBertolani et al. 2023Macrobiotus siderophilusC3282_UFOOP561781Macrobiotus siderophilusBertolani et al. 2023Macrobiotus siderophilusC3282_UFPOP561782Macrobiotus siderophilusBertolani et al. 2023Macrobiotus sottileiPL.352MW246133MW247178MW247179MW247175Macrobiotus sottileiKiosya et al. 2021Macrobiotus sp.DI02OP684780OP696675Macrobiotus sp.Amezcua-Martinez et al. (unpubl.)Macrobiotus sp. 1 MC-2022dUK 12OQ525891OQ525899OQ548104OQ525904Macrobiotus sp. 1 MC-2022dMioduchowsk et al. (unpubl.)Macrobiotus sp. 1 MC-2022dUK 14OQ525890OQ525897OQ525905Macrobiotus sp. 1 MC-2022dMioduchowsk et al. (unpubl.)Macrobiotus sp. 1 YW-2020BN140005MT863704MT872214Macrobiotus sp. 1 YW-2020Wang (unpubl.)Macrobiotus sp. 1 ZY-2020HP1110201MW187003MW183923Macrobiotus sp. 1 ZY-2020Yuan et al. (unpubl.)Macrobiotus sp. n. MM-2020MAD967MT246661MT241900MT241898Macrobiotus sp. n. MM-2020Kuzdrowska et al. (unpubl.)Macrobiotus terminalisC2868_N02JN673959OP596308Macrobiotus terminalisCesari et al. 2011Macrobiotus vattenrikense sp. nov22-003PX093655PX093663PX093644PX093649COI haplotype 3; 18S haplotype 1; ITS haplotype 1; 28S haplotype 1this studyMacrobiotus vattenrikense sp. nov22-004-1PX093655COI haplotype 3this studyMacrobiotus vattenrikense sp. nov22-017PX093653PX093644PX093649COI haplotype 1; ITS haplotype 1; 28S haplotype 1this studyMacrobiotus vattenrikense sp. nov22-101PX093654PX093664PX093644PX093650COI haplotype 2; 18S haplotype 2; ITS haplotype 1; 28S haplotype 2this studyMacrobiotus vladimiriFI.068MN888327MN888375MN888347MN888360Macrobiotus vladimiriVecchi & Stec 2021Mesobiotus aradasiKPRI-MF-1MW727934MW751962Mesobiotus cf. furcigersee Vecchi et al. 2024aShort et al. 2022Mesobiotus aradasiS1982.Meb.1OQ928737OQ933000OQ940183OQ932999Mesobiotus aradasiVecchi et al. 2024aMesobiotus bockebodicus22-044PQ365775PQ367888PQ367894Mesobiotus sp. 1 SA-2024atemporary taxon name has not been updated in GenBank at time of writingAtherton et al. 2025Mesobiotus cf. barabanoviKG.002MN313170MN310392MN310390MN310388Mesobiotus cf. barabanoviKaczmarek et al. 2020aMesobiotus cf. furcigerAF01-MF-2MW727944MW751946Mesobiotus cf. furcigerShort et al. 2022Mesobiotus datanlanicusVN.026MK578905MK584659MK584657MK584658Mesobiotus datanlanicusStec 2019Mesobiotus diegoiZA.001.02OP143857OP142526OP142514OP142520Mesobiotus diegoiStec 2022aMesobiotus dilimanensisPH.006MN257047MN257048MN257050MN257049Mesobiotus dilimanensisItang et al. 2020Mesobiotus efaLembolovo.DT262OR803035OR804457OR805169OR805135Mesobiotus sp. 1 DVT-2023btemporary taxon name has not been updated in GenBank at time of writingTumanov et al. 2024Mesobiotus emiliae22-046PQ365768PQ367886PQ367892Mesobiotus emiliaeAtherton et al. 2025Mesobiotus ethiopicusET.004MF678794MF678793MN122776MF678792Mesobiotus ethiopicusStec & Kristensen 2017Mesobiotus fiedleriMG.001MH676056MH681585MH681724MH681693Mesobiotus fiedleriKaczmarek et al. 2020aMesobiotus furcigerMacro_06_313MW727960MW751939Mesobiotus furcigerShort et al. 2022Mesobiotus harmsworthihaplotype 1MH195150MH197146MH197154MH197264Mesobiotus harmsworthiKaczmarek et al. 2018Mesobiotus hilariaeNovolazarevskaya_ C_1KT226108KT226071Mesobiotus hilariaeVecchi et al. 2016Mesobiotus huecoensisS2027.4OQ756247OQ756248Mesobiotus huecoensisVecchi et al. 2023Mesobiotus imperialisVN.061.01OL311514OL257855OL257867Mesobiotus imperialisStec 2021Mesobiotus insanisMF 441491MF441488MF441490MF441489Mesobiotus insanisMapalo et al. 2017Mesobiotus maklowicziZA.002.01OP143855OP142525OP142518Mesobiotus maklowicziStec 2022aMesobiotus mandalori2.18.1.2OP825090OP829143OP829058OP829056Mesobiotus sp. m MM-2022temporary taxon name has not been updated in GenBank at time of writingErdmann et al. 2024Mesobiotus mandalori22-053PQ365769PQ367887PQ367893Mesobiotus sp. m MM-2022temporary taxon name has not been updated in GenBank at time of writingAtherton et al. 2025Mesobiotus marmoreusVN.055.01OL311516OL257856OL257861OL257868Mesobiotus marmoreusStec 2022aMesobiotus occultatusDT97OR803042OR794157OR805249OR794158Mesobiotus occultatusTumanov et al. 2024Mesobiotus peterseniGL.002.01OP143859OP142528OP142516OP142522Mesobiotus peterseniStec 2022aMesobiotus philippinicusKX 129796KX129793KX129795KX129794Mesobiotus philippinicusMapalo et al. 2016Mesobiotus radiatusKE.008MH195147MH197153MH197267MH197152Mesobiotus radiatusStec et al. 2018dMesobiotus romaniEC.002MH195149MH197158MH197150MH197151Mesobiotus romaniRoszkowska et al. 2018Mesobiotus skanensis22-153PQ365777PQ367889PQ367895Mesobiotus sp. 2 SA-2024atemporary taxon name has not been updated in GenBank at time of writingAtherton et al. 2025Mesobiotus skorackiiCN8MW656257MW680643MW680636Mesobiotus skorackiiKayastha et al. 2021Mesobiotus sp.BI02OP684777OP696672Mesobiotus sp.Amezcua-Martinez et al. (unpubl.)Mesobiotus sp.NO.234MH195153MH197148MH197156MH197265Mesobiotus furciger group sp.Kaczmarek et al. 2018Mesobiotus sp.RU.017MH195154MH197149MH197157MH197266Mesobiotus harmsworthi group sp.Kaczmarek et al. 2018Mesobiotus sp. 1DVT-2020MT904513MT903468MT903470MT903612Mesobiotus sp. 1Tumanov 2020Mesobiotus sp. FlMes_sp_S16_3OQ968323OQ974689OQ974696OQ974704Mesobiotus sp. FlVecchi et al. 2024bMesobiotus sp. VNMes_sp_ VN 036_04OQ968314OQ974691Mesobiotus sp. VNVecchi et al. 2024bMesobiotus vulpinusDT553OR803040OR804461OR805172OR805140Mesobiotus sp. 2 DVT-2023btemporary taxon name has not been updated in GenBank at time of writingTumanov et al. 2024Mesobiotus zelmae22-239PQ365781PQ367891PQ367896Mesobiotus sp. 3 SA-2024atemporary taxon name has not been updated in GenBank at time of writingAtherton et al. 2025Mesobiotus cf. intermediusDT270OP013287OP035718OP035707OP035798Minibiotus cf. intermediusTumanov et al. 2022Minibiotus citlaliumAI04OP684765OP696660Minibiotus citlaliumAmezcua-Martinez et al. (unpubl.)Minibiotus furcatusTar527FJ435802FJ435746FJ435760Minibiotus furcatusGuil & Giribet 2012Minibiotus gumersindoiTar710FJ435803FJ435748FJ435761Minibiotus gumersindoiGuil & Giribet 2012Minibiotus intermediusMin4 GRON005160ON005189ON005195Minibiotus intermediusKaczmarek et al. 2022Minibiotus ioculatorZA.274MT023412MT023998MT024000MT024041Minibiotus ioculatorStec et al. 2020bMinibiotus pentannulatusTZ.027 haplotype 1 MT023413MT023999MT024001MT024042Minibiotus pentannulatusStec et al. 2020bMinibiotus sidereusAI12OP684770OP696665Minibiotus sidereusAmezcua-Martinez et al. (unpubl.)Minibiotus sp. A MV2020S69.Min.01MW306859OK663227OK663216OK663238Minibiotus sp. A MV2020Vecchi et al. 2022bParamacrobiotus marchelmoni sp. nov.22-004PX093656PX093665PX093646PX093651COI haplotype 1; 18S haplotype 1; ITS haplotype 2; 28S haplotype 1this studyParamacrobiotus marchelmoni sp. nov.22-006PX093656PX093665PX093646COI haplotype 1; 18S haplotype 1; ITS haplotype 2this studyParamacrobiotus marchelmoni sp. nov.22-007PX093656PX093646COI haplotype 1; ITS haplotype 2this studyParamacrobiotus marchelmoni sp. nov.22-019PX093656PX093665PX093646PX093651COI haplotype 1; 18S haplotype 1; ITS haplotype 2; 28S haplotype 1this studyParamacrobiotus marchelmoni sp. nov.22-026PX093656COI haplotype 1this studyParamacrobiotus marchelmoni sp. nov.22-029PX093656PX093645PX093651COI haplotype 1; 18S haplotype 1; 28S haplotype 1this studyParamacrobiotus marchelmoni sp. nov.22-034PX093656PX093665PX093646COI haplotype 1; 18S haplotype 1; ITS haplotype 2this studyParamacrobiotus marchelmoni sp. nov.22-102PX093656PX093645COI haplotype 1; ITS haplotype 1this studyParamacrobiotus marchelmoni sp. nov.22-106PX093657COI haplotype 2this studyParamacrobiotus marchelmoni sp. nov.22-108PX093656COI haplotype 1this studyParamacrobiotus marchelmoni sp. nov.22-110PX093657PX093665PX093646PX093651COI haplotype 2; 18S haplotype 1; ITS haplotype 2; 28S haplotype 1this studyParamacrobiotus marchelmoni sp. nov.22-113PX093657PX093665PX093646COI haplotype 2; 18S haplotype 1; ITS haplotype 2this studyParamacrobiotus marchelmoni sp. nov.23-015PX093658COI haplotype 3this studyParamacrobiotus marchelmoni sp. nov.23-041PX093657COI haplotype 2this studyParamacrobiotus aff. arduusPRODO-3MK041022Paramacrobiotus aff. arduusGuidetti et al. 2019aParamacrobiotus aff. spatialis OLBIA-1MK041002MK041026Paramacrobiotus aff. spatialisGuidetti et al. 2019aParamacrobiotus arduusPRODO-1MK041020MK041032Paramacrobiotus arduusGuidetti et al. 2019aParamacrobiotus arduusPRODO-2MK041021Paramacrobiotus arduusGuidetti et al. 2019aParamacrobiotus areolatusNO.385MH675998MH664931MH666080MH664948Paramacrobiotus areolatusStec et al. 2020cParamacrobiotus bifronsC4497_ US 1PP236542PP240910Paramacrobiotus sp. 1 MC-2024atemporary taxon name has not been updated in GenBank at time of writingBrandoli et al. 2024Paramacrobiotus bifronsC4497_ US 2PP236543PP240911Paramacrobiotus sp. 1 MC-2024atemporary taxon name has not been updated in GenBank at time of writingBrandoli et al. 2024Paramacrobiotus bifronsC4497_ US 7PP236544PP240912Paramacrobiotus sp. 1 MC-2024atemporary taxon name has not been updated in GenBank at time of writingBrandoli et al. 2024Paramacrobiotus bifronsC4497_ US 8PP236545PP240913Paramacrobiotus sp. 1 MC-2024atemporary taxon name has not been updated in GenBank at time of writingBrandoli et al. 2024Paramacrobiotus bifronsC5025_ US 6PP236551PP240918Paramacrobiotus sp. 1 MC-2024atemporary taxon name has not been updated in GenBank at time of writingBrandoli et al. 2024Paramacrobiotus bifronsC5062_ US 2PP236553PP240919Paramacrobiotus sp. 1 MC-2024atemporary taxon name has not been updated in GenBank at time of writingBrandoli et al. 2024Paramacrobiotus bifronsC5062_ US 3PP236554PP240920Paramacrobiotus sp. 1 MC-2024atemporary taxon name has not been updated in GenBank at time of writingBrandoli et al. 2024Paramacrobiotus bifronsIT.048MH676007MH664937MH666085MH664955Paramacrobiotus sp.see Brandoli et al. 2024Stec et al. 2020cParamacrobiotus bifronsPar_cf_kly_ S108_2OQ968322OQ974697Paramacrobiotus cf. klymenkisee Brandoli et al. 2024Vecchi et al. 2024bParamacrobiotus celsusPIANE_DI_ MOCOGNO-1MK041017MK041031Paramacrobiotus celsusGuidetti et al. 2019aParamacrobiotus celsusPIANE_DI_ MOCOGNO-2MK041018Paramacrobiotus celsusGuidetti et al. 2019aParamacrobiotus celsusPIANE_DI_ MOCOGNO-3MK041019Paramacrobiotus celsusGuidetti et al. 2019aParamacrobiotus danielaeGY26001MZ081363MZ081364Paramacrobiotus danielaeWang et al. (unpubl.)Paramacrobiotus depressusANDALO-1MK041015Paramacrobiotus depressusGuidetti et al. 2019aParamacrobiotus depressusOSPITALETTO_ A-1MK041016Paramacrobiotus depressusGuidetti et al. 2019aParamacrobiotus depressusPASSO_ BALLINO-1MK041012MK041030Paramacrobiotus depressusGuidetti et al. 2019aParamacrobiotus depressusPASSO_ BALLINO-2MK041013Paramacrobiotus depressusGuidetti et al. 2019aParamacrobiotus depressusPASSO_ BALLINO-3MK041014Paramacrobiotus depressusGuidetti et al. 2019aParamacrobiotus experimentalisMAD-TAR11MN097837MN073464MN073465Paramacrobiotus experimentalisKaczmarek et al. 2020cParamacrobiotus fairbanksi22-423PX093660PX093666Paramacrobiotus fairbanksithis studyParamacrobiotus fairbanksi23-022PX093661Paramacrobiotus fairbanksithis studyParamacrobiotus fairbanksi23-041PX093659PX093666PX093647PX093648Paramacrobiotus fairbanksithis studyParamacrobiotus fairbanksiAlb1ON911918Paramacrobiotus fairbanksiMioduchowsk et al. (unpubl.)Paramacrobiotus fairbanksiAlb4ON911917Paramacrobiotus fairbanksiMioduchowsk et al. (unpubl.)Paramacrobiotus fairbanksiC2345-MR_2AY598778Macrobiotus richtersiGuidetti et al. 2005Paramacrobiotus fairbanksiC2345-MR_3AY598779Macrobiotus richtersiGuidetti et al. 2005Paramacrobiotus sp.Kenya 3EU244599Paramacrobiotus sp. ‘ richtersi ’ groupSchill et al. 2010Paramacrobiotus sp.MG.002MH676008MH664938MH666086MH664956Paramacrobiotus sp.Stec et al. 2020cParamacrobiotus sp.NO.386MH676009MH664939MH666088MH664957Paramacrobiotus sp.Stec et al. 2020cParamacrobiotus sp.NZ.001MH676010MH664940MH666089MH664958Paramacrobiotus sp.Stec et al. 2020cParamacrobiotus sp.Par_sp_S175_2OQ968321Paramacrobiotus sp.Vecchi et al. 2024bParamacrobiotus sp.PT.006MH676013MH664943MH666092MH664960Paramacrobiotus sp.Stec et al. 2020cParamacrobiotus sp.PT.048 haplotype 1MH676014MH664944MH666093MH664961Paramacrobiotus sp.Stec et al. 2020cParamacrobiotus sp.PT.048 haplotype 2MH676015Paramacrobiotus sp.Stec et al. 2020cParamacrobiotus sp.T3OP531839ON923868Paramacrobiotus sp.Basu et al. 2023Paramacrobiotus sp.Tar407FJ435807FJ435741FJ435756Macrobiotus pallariisee DiscussionGuil & Giribet 2012Paramacrobiotus sp.TN.014MH676016MH664945MH666094MH664962Paramacrobiotus sp.Stec et al. 2020cParamacrobiotus sp.TZ.018MH676017MH664933MH666095MH664951Paramacrobiotus sp.Stec et al. 2020cParamacrobiotus sp. DLS-2024aT82_S135PP414782PP416751PP416753PP416752Paramacrobiotus sp. DLS-2024aLópez-Sandoval et al. 2025Paramacrobiotus sp. MAC-2014T1KF788251Paramacrobiotus richtersi group sp. 1 MAC-2014Caicedo et al. 2017Paramacrobiotus sp. MAC-2014T2KF788252Paramacrobiotus richtersi group sp. 1 MAC-2014Caicedo et al. 2017Paramacrobiotus sp. MAC-2014T3KF788253Paramacrobiotus richtersi group sp. 1 MAC-2014Caicedo et al. 2017Paramacrobiotus sp. MAC-2014T4KF788254Paramacrobiotus richtersi group sp. 1 MAC-2014Caicedo et al. 2017Paramacrobiotus sp. MAC-2014T6KF788255Paramacrobiotus richtersi group sp. 1 MAC-2014Caicedo et al. 2017Paramacrobiotus sp. MAC-2014T7KF788256Paramacrobiotus richtersi group sp. 1 MAC-2014Caicedo et al. 2017Paramacrobiotus sp. MAC-2014T8KF788257Paramacrobiotus richtersi group sp. 1 MAC-2014Caicedo et al. 2017Paramacrobiotus sp. ZY-2020hp1100404MT731035MT723894Paramacrobiotus sp. ZY-2020Yuan et al. (unpubl.)Paramacrobiotus sp. ZY-2022cN 3OP870172ON819001Paramacrobiotus sp. ZY-2022cYuan et al. (unpubl.)Paramacrobiotus spatialisFORMIGINE-1MK040995MK041024Paramacrobiotus spatialisGuidetti et al. 2009Paramacrobiotus spatialisFORMIGINE-2MK040996Paramacrobiotus spatialisGuidetti et al. 2019aParamacrobiotus spatialisFORMIGINE-3MK040997Paramacrobiotus spatialisGuidetti et al. 2019aParamacrobiotus spatialisFORMIGINE-4MK040998Paramacrobiotus spatialisGuidetti et al. 2019aParamacrobiotus spatialisGAGGIO-1MK040999Paramacrobiotus spatialisGuidetti et al. 2019aParamacrobiotus spatialisGAGGIO-2MK041000Paramacrobiotus spatialisGuidetti et al. 2019aParamacrobiotus spatialisGAGGIO-3MK041001MK041025Paramacrobiotus spatialisGuidetti et al. 2019aParamacrobiotus spatialisS107.ParOK662996OK663225OK663214OK663236Paramacrobiotus spatialisVecchi et al. 2022cParamacrobiotus tonolliiUS.035MH676018MH664946MH666096MH664963Paramacrobiotus tonolliiStec et al. 2020cRichtersius coroniferNO.385MH676053MH681760MH681763MH681757Richtersius coroniferStec et al. 2020cSisubiotus hakaiensisSis.hak.S1911OM523181OM523054OM523057OM523059Sisubiotus hakaiensisVecchi et al. 2022aSisubiotus spectabilisFI.067MN888322MN888371MN888331MN888357Sisubiotus spectabilisStec et al. 2021aTenuibiotus cf. ciprianoiES.086MN888328MN888376MN888348MN888361Tenuibiotus cf. ciprianoiStec et al. 2021aTenuibiotus daniloviKG.128MN888329MN888377MN888349MN888362Tenuibiotus daniloviStec et al. 2021aTenuibiotus sp.22-109PX093662PX093667PX093652Tenuibiotus sp.this studyTenuibiotus tenuiformisKG.140MN888330MN888378MN888350MN888363Tenuibiotus tenuiformisStec et al. 2021aTenuibiotus voronkovihaplotype 1KX810042KX810045KX810046KX810049Tenuibiotus voronkoviZawierucha et al. 2016Tenuibiotus yeliseiiDT422OR145334OR142418OR142424OR142426Tenuibiotus sp. 1 DVT-2023atemporary taxon name has not been updated in GenBank at time of writingTsvetkova & Tumanov 2024Tenuibiotus zandraeMN 444827MN443040MN443038MN443035Tenuibiotus zandraeStec et al. 2020cXerobiotus arenosumC4411_V4OR397039OR398024OR398050Xerobiotus arenosumVincenzi et al. 2024Xerobiotus cf. reductusPL3177_1OR397050OR398033OR398056Xerobiotus cf. reductusVincenzi et al. 2024Xerobiotus euxinuscheck 1.4PP469626PP439986PP470593PP440001Macrobiotus euxinussee Vincenzi et al. 2024Polishchuk et al. 2024Xerobiotus euxinusC3834_V4OR397005OR398005OR398119Macrobiotus euxinussee Vincenzi et al. 2024Vincenzi et al. 2024Xerobiotus gretaeC4341 GMW581665MW588434MW588431MW588438Macrobiotus gretaesee Vincenzi et al. 2024Massa et al. 2021Xerobiotus litusC4383_2OR397000OR398002OR398118Xerobiotus litusVincenzi et al. 2024Xerobiotus naginaeS226.Mac.1OK662990OK663219OK663230Marobiotus naginaesee Vincenzi et al. 2024Vecchi et al. 2022cXerobiotus pseudohufelandiC2358-XP1AY598776HQ604989Xerobiotus pseudohufelandiGuidetti et al. 2005Xerobiotus reductusPL_3176_1OR397048OR398028OR398130Xerobiotus reductusVincenzi et al. 2024Xerobiotus sp.PL.360MN888325MN888373MN888345MN888358Xerobiotus sp.Vecchi & Stec 2021Xerobiotus sp.ZA.373MN888326MN888374MN888346MN888359Xerobiotus sp.Vecchi & Stec 2021Xerobiotus sp. 1 MC-2023aC3306_V1OR397025OR398020OR398123Xerobiotus sp. 1 MC-2023aVincenzi et al. 2024