Table 2 in Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data

Table 2. Sampling of Nebriitae and outgroup species. The ID column indicates how each voucher specimen was identified. A number indicates that the specimen was identified by the lead author (DK) using the corresponding numbered references as follows: (1) Lindroth (1961), (2) Andrewes (1929), (3) Bousquet and Smetana (1996), (4) Kasahara (1989), (5) Uéno (1955), (6) Perrault (1981), (8) Jeannel (1941), (9) Perrault (1985), (10) Minowa (1932), (11) Farkač (1995), (12) Perrault (1986), (13) Erwin (1970), (14) Dudko (2003), (15) Perrault (1991), (16) Huber and Schmidt (2016), (17) Ledoux and Roux (2005), (18) Shilenkov (1975), (19) Ledoux and Roux (2009), (20) Trautner (1987), (21) Huber and Schmidt (2013), (22) Dudko (2006) and (23) Sasakawa (2020). A “T” indicates that the specimen was compared with the primary type specimen. A two-letter code indicates that the specimen was identified by one of the following: AF = A. Faille; DM = D.R. Maddison; JS = J. Schmidt; and RD = R.Y. Dudko. Three-letter codes indicate the following: CRC = compared with material in Philippe Roux collection and matching no taxon therein; and NID = could not be identified to species. In the remaining columns, GenBank numbers (MW359101 through MW361062) are provided for each specimen from which the listed gene sequence was successfully recovered in this study. Additional information on these voucher specimens is provided in Appendix A. Other GenBank numbers are for previously published sequences from Kavanaugh et al. (2011), Maddison (2008), Maddison et al. (2009), Ober (2002), and Wild and Maddison (2008). Sequences of the five specimens marked with an asterisk in the ID column are paratypes and the sequences are thus “genseq-2”, with the remainder being “genseq-4” (Chakrabarty et al. 2013).
ID28S16S-ND1CO1 BCCO1 PJCAD2PEPCKTopowgSupertribe NebriitaeTribe NotiophiliniGenus Notiophilus DumérilNotiophilus borealis Harris1MW359101MW360814MW359348MW359611MW360101MW360345MW359863MW360574Notiophilus semistriatus Say1MW359102MW360815MW359349MW359612MW360102MW360346MW359864MW360575Notiophilus sierranus Casey1MW359103MW360816MW359350MW359613MW360103MW360347MW359865MW360576Notiophilus sylvaticus Dejean1MW359104MW360817MW359351MW359614MW360104MW360348MW359866MW360577Tribe OpisthiiniGenus Opisthius KirbyOpisthius richardsoni Kirby1MW359105MW360818MW359352MW359615MW360105MW360349MW359867MW360578Genus Paropisthius CaseyParopisthius davidis (Fairmaire)2MW359106MW360819MW359353MW359616MW360106MW360350MW359868MW360579Paropisthius indicus chinensis Bousquet & Smetana3MW359107MW360820MW359354MW359617MW360107MW360351MW359869MW360580Paropisthius indicus indicus (Chaudoir)3MW359108MW360821MW359355MW359618MW360108MW360352MW359870MW360581Paropisthius masuzoi Kasahara4MW359109MW360822MW359356MW359619MW360109MW360353MW359871MW360582Tribe PelophiliniGenus Pelophila DejeanPelophila borealis (Paykull) AK1MW359110MW360823MW359357MW359620MW360110MW360354MW359872MW360583Pelophila borealis (Paykull) RFE1MW359111-MW359358MW359621MW360111-MW359873MW360584Pelophila rudis (LeConte)1MW359112MW360824MW359359MW359622MW360112MW360355MW359874MW360585Tribe NebriiniGenus Nippononebria UénoSubgenus Nippononebria Nippononebria UénoNippononebria chalceola (Bates)5--MN982525MN982525----Nippononebria changbaiensis Kavanaugh & LiangTMW359113MW360825MW359360MW359623MW360113MW360356MW359875MW360586Subgenus Nippononebria Vancouveria KavanaughNippononebria altisierrae KavanaughTMW359114MW360826MW359361MW359624MW360114MW360357MW359876MW360587Nippononebria campbelli KavanaughTMW359115MW360827MW359362MW359625MW360115MW360358MW359877MW360588Nippononebria virescens (Horn)TMW359116MW360828MW359363MW359626MW360116MW360359MW359878MW360589Genus Leistus FrölichSubgenus Leistus Nebrileistus BänningerLeistus nubivagus Wollaston6MW359117MW360829MW359364MW359627MW360117MW360360MW359879MW360590Subgenus Leistus Sardoleistus PerraultLeistus sardous Baudi di SelveAFMW359118MW360830MW359365MW359628MW360118--MW360591Subgenus Leistus Pogonophorus LatreilleLeistus parvicollis Chaudoir8MW359119MW360831MW359366MW359629MW360119MW360361-MW360592Leistus pyrenaeus Kraatz8MW359120MW360832MW359367MW359630MW360120-MW359880MW360593Leistus rufomarginatus (Duftschmid)8MW359121MW360833MW359368MW359631MW360121MW360362MW359881MW360594Subgenus Leistus Evanoleistus JedličkaLeistus birmanicus Perrault9MW359122MW360834MW359369MW359632MW360122MW360363MW359882MW360595Leistus gaoligongensis Kavanaugh & LongTMW359123MW360835MW359370MW359633MW360123MW360364MW359883MW360596Leistus lihengae Kavanaugh & LongTMW359124MW360836MW359371MW359634MW360124MW360365MW359884MW360597Leistus niitakaensis Minowa10MW359125MW360837MW359372MW359635MW360125MW360366MW359885MW360598Leistus nokoensis Minowa10MW359126MW360838MW359373MW359636MW360126MW360367MW359886MW360599Leistus smetanai Farkač11MW359127MW360839MW359374MW359637MW360127MW360368MW359887MW360600Leistus taiwanensis Perrault12MW359128MW360840MW359375MW359638MW360128MW360369MW359888MW360601Leistus sp NEPNIDMW359129MW360841MW359376MW359639MW360129-MW359889MW360602Leistus sp YUN 1NIDMW359130MW360842MW359377MW359640MW360130MW360370MW359890MW360603Subgenus Leistus Leistus FrölichLeistus crenatus Fairmaire8MW359131MW360843MW359378MW359641MW360131MW360371MW359891MW360604Leistus ferrugineus (Linnaeus)8MW359132MW360844MW359379MW359642MW360132MW360372MW359892MW360605Leistus ferruginosus Mannerheim13MW359133MW360845MW359380MW359643MW360133MW360373MW359893MW360606Leistus kryzhanovskii frateroides Dudko14MW359134MW360846MW359381MW359644MW360134MW360374MW359894MW360607Leistus kryzhanovskii kryzhanovskii Dudko14MW359135MW360847MW359382MW359645MW360135MW360375MW359895MW360608Leistus fulvibarbis Dejean8MW359136MW360848MW359383MW359646MW360136--MW360609Leistus fulvus Chaudoir8-MW360849MW359384MW359647----Leistus longipennis Casey13MW359137MW360850MW359385MW359648MW360137MW360376MW359896MW360610Leistus madmeridianus Erwin13MW359138MW360851MW359386MW359649MW360138MW360377MW359897MW360611Leistus niger Gebler15MW359139MW360852MW359387MW359650MW360139MW360378MW359898MW360612Leistus nitidus (Duftschmid)8MW359140MW360853MW359388MW359651MW360140MW360379-MW360613Leistus sp JIL 2NIDMW359141MW360854MW359389MW359652MW360141MW360380MW359899MW360614Leistus sp SCHNIDMW359142MW360855MW359390MW359653MW360142MW360381MW359900MW360615Leistus sp YUN 2NIDMW359143MW360856MW359391MW359654MW360143MW360382MW359901MW360616Genus Nebria LatreilleOreonebria SeriesEonebria ComplexSubgenus Eonebria Pseudepinebriola New SubgenusNebria delicata Huber & Schmidt116MW359144MW360857MW359392MW359655MW360144MW360383MW359902MW360617Nebria retingensis Huber & Schmidt116*MW359145MW360858MW359393MW359656MW360145MW360384MW359903MW360618Nebria sp YUN 4NIDMW359146MW360859MW359394MW359657MW360146MW360385MW359904MW360619Nebria sp YUN 5NIDMW359147MW360860MW359395MW359658MW360147MW360386MW359905MW360620Nebria sp XIZ 14NIDMW359148MW360861MW359396MW359659MW360148MW360387MW359906MW360621Subgenus Eonebria Sadonebria Ledoux & RouxNebria chinensis Bates17MW359149MW360862MW359397MW359660MW360149MW360388MW359907MW360622Nebria niitakana Kano17MW359150MW360863MW359398MW359661MW360150MW360389MW359908MW360623Nebria ohdaiensis Nakane17MW359151MW360864MW359399MW359662MW360151MW360390MW359909MW360624Nebria saeviens Bates17MW359152MW360865MW359400MW359663MW360152MW360391MW359910MW360625Subgenus Eonebria Eonebria Semenov & ZnojkoNebria komarovi Semenov & Znojko17MW359153MW360866MW359401MW359664MW360153-MW359911-Nebria sp GANCRCMW359154MW360867MW359402MW359665MW360154-MW359912-Nebria sp YUN 13CRCMW359155MW360868MW359403MW359666MW360155MW360392MW359913MW360626Nebria sp YUN 16CRCMW359156MW360869MW359404MW359667MW360156MW360393MW359914MW360627Nebria sp YUN 17CRCMW359157MW360870MW359405MW359668MW360157MW360394MW359915MW360628Nebria sp YUN 18CRCMW359158MW360871MW359406MW359669MW360158MW360395MW359916MW360629Oreonebria ComplexSubgenus Oreonebria Epispadias Ledoux & RouxNebria sp YUN 6CRCMW359159MW360872MW359407MW359670MW360159MW360396MW359917MW360630Subgenus Oreonebria Falcinebria Ledoux & RouxNebria formosana Habu17MW359160MW360873MW359408MW359671MW360160MW360397MW359918MW360631Nebria niohozana Bates23MW359161MW360874MW359409MW359672MW360161MW360398MW359919MW360632Subgenus Nebria Orientonebria ShilenkovNebria coreica Solsky18MW359162MW360875MW359410MW359673MW360162MW360399MW359920MW360633Subgenus Nebria Archastes JedličkaNebria yuae (Ledoux & Roux)19MW359163MW360876MW359411MW359674MW360163MW360400MW359921MW360634Subgenus Nebria Oreonebria DanielNebria angustata Dejean17MW359164MW360877MW359412MW359675MW360164MW360401MW359922MW360635Nebria angusticollis Bonelli17MW359165MW360878MW359413MW359676MW360165MW360402MW359923MW360636Nebria austriaca Ganglbauer17MW359166MW360879MW359414MW359677MW360166-MW359924MW360637Nebria boschi Winkler17MW359167MW360880MW359415MW359678MW360167MW360403MW359925MW360638Nebria bremii Germar217MW359168MW360881MW359416MW359679MW360168MW360404MW359926MW360639Nebria castanea Bonelli17MW359169MW360882MW359417MW359680MW360169MW360405MW359927MW360640Nebria diaphana Daniel & Daniel17MW359170MW360883MW359418MW359681MW360170-MW359928MW360641Nebria gagates (Bonelli)317MW359171MW360884MW359419MW359682MW360171-MW359929MW360642Nebria ligurica Daniel17MW359172MW360885MW359420MW359683MW360172-MW359930MW360643Nebria lombarda Daniel & Daniel17MW359173-MW359421MW359684MW360173-MW359931MW360644Nebria picea Dejean17MW359174MW360886MW359422MW359685MW360174MW360406MW359932MW360645Nebriola SeriesSubgenus Nebria Nebriola DanielNebria fontinalis Daniel & Daniel17MW359175MW360887MW359423MW359686MW360175MW360407MW359933MW360646Nebria laticollis Dejean17MW359176MW360888MW359424MW359687MW360176MW360408MW359934MW360647Nebria SeriesBoreonebria ComplexSubgenus Nebria Nakanebria Ledoux & RouxNebria shiretokoana Nakane17MW359177MW360889MW359425MW359688MW360177MW360409MW359935MW360648Subgenus Nebria Boreonebria JeannelNebria baicalica Motschulsky18JN847585JN847610MW359426JN847535JN847560MW360410JN847510JN847635Nebria bellorum KavanaughTJN847603JN847628MW359427JN847553JN847578MW360411JN847528JN847653Nebria castanipes KirbyTMW359178MW360890MW359428MW359689MW360178MW360412MW359936MW360649Nebria changaica Horvatovich17MW359179MW360891MW359429MW359690MW360179-MW359937MW360650Nebria crassicornis Van Dyke1JN847581JN847606MW359430JN847531JN847556-JN847506JN847631Nebria dabanensis Shilenkov417MW359180MW360892MW359431MW359691MW360180-MW359938MW360651Nebria frigida Sahlberg1MW359181MW360893MW359432MW359692MW360181MW360413MW359939MW360652Nebria gouleti KavanaughTJN847587JN847612MW359433JN847537JN847562MW360414JN847512JN847637Nebria gyllenhali (Schönherr)TJN847582JN847607MW359434JN847532JN847557-JN847507JN847632Nebria hudsonica LeConteTJN847589JN847614MW359435JN847539JN847564MW360415JN847514JN847639Nebria intermedia Van Dyke OR1MW359182MW360894MW359436MW359693MW360182-MW359940MW360653Nebria intermedia Van Dyke UT1MW359183MW360895MW359437MW359694MW360183-MW359941MW360654Nebria intermedia Van Dyke WY1MW359184MW360896MW359438MW359695MW360184-MW359942MW360655Nebria kaszabi Shilenkov417MW359185MW360897MW359439MW359696MW360185-MW359943MW360656Nebria lacustris CaseyTJN847590JN847615MW359440JN847540JN847565MW360416JN847515JN847640Nebria lassenensis KavanaughTMW359186MW360898MW359441MW359697MW360186MW360417MW359944MW360657Nebria lindrothi KavanaughTMW359187MW360899MW359442MW359698MW360187MW360418MW359945MW360658Nebria murzini Ledoux & Roux417MW359188MW360900MW359443MW359699MW360188-MW359946MW360659Nebria nivalis (Paykull) AK1JN847584JN847609MW359444JN847534JN847559MW360419JN847509JN847634Nebria nivalis (Paykull) NU1JN847583JN847608MW359445JN847533JN847558MW360420JN847508JN847633Nebria nivalis gr spNIDMW359189MW360901MW359446MW359700MW360189MW360421MW359947MW360660Nebria rubrofemorata Shilenkov18MW359190MW360902MW359447MW359701MW360190MW360422MW359948MW360661Nebria sajanica Bänninger 417MW359191MW360903MW359448MW359702MW360191-MW359949MW360662Nebria sajanica gr sp 14NIDMW359192MW360904MW359449MW359703---MW360663Nebria sajanica gr sp 24NIDMW359193MW360905MW359450MW359704MW360192MW360423MW359950MW360664Nebria subdilatata Motschulsky18JN847586JN847611MW359451JN847536JN847561MW360424JN847511JN847636Nebria tekesensis Ledoux & Roux417MW359194MW360906MW359452MW359705MW360193-MW359951MW360665Nebria sp MG 1NIDMW359195MW360907MW359453MW359706MW360194MW360425MW359952MW360666Nebria sp MG 2NIDMW359196MW360908MW359454MW359707MW360195MW360426MW359953MW360667Nebria ComplexNebria SubcomplexSubgenus Nebria Tyrrhenia Ledoux & RouxNebria fulviventris Bassi17MW359197MW360909MW359455MW359708MW360196MW360427MW359954MW360668Nebria rubicunda maroccana Antoine17MW359198MW360910MW359456MW359709MW360197-MW359955MW360669Subgenus Nebria Nebria LatreilleNebria brevicollis (Fabricius) OR20JN847580MW360911MW359457JN847530JN847555MW360428JN847505JN847630Nebria brevicollis (Fabricius) SP20MW359199MW360912MW359458MW359710MW360198MW360429MW359956MW360670Nebria lafresnayi Audinet-Serville8MW359200MW360913MW359459MW359711MW360199-MW359957MW360671Nebria olivieri Dejean8MW359201MW360914MW359460MW359712MW360200MW360430MW359958MW360672Nebria rubripes Audinet-Serville8MW359202MW360915MW359461MW359713MW360201MW360431MW359959MW360673Nebria tibialis Bonelli17MW359203MW360916MW359462MW359714MW360202MW360432MW359960MW360674Nebria turcica Chaudoir17MW359204MW360917MW359463MW359715MW360203--MW360675Subgenus Nebria Alpaeonebria CsikiNebria germari Heer AU17MW359205MW360918MW359464MW359716MW360204MW360433MW359961MW360676Nebria germari Heer GE17MW359206MW360919MW359465MW359717MW360205MW360434MW359962MW360677Subgenus Nebria Spelaeonebria PeyerimhoffNebria nudicollis PeyerimhoffAFMW359207MW360920MW359466MW359718MW360206-MW359963MW360678Epinebriola SubcomplexSubgenus Nebria Epinebriola DanielNebria businskyorum Ledoux & Roux16MW359208MW360921MW359467MW359719MW360207MW360435MW359964MW360679Nebria capillosa Ledoux & Roux6JSMW359209MW360922MW359468MW359720MW360208MW360436MW359965-Nebria cf. desgodinsi Oberthür 6JSMW359210MW360923MW359469MW359721MW360209-MW359966-Nebria kagmara Huber & Schmidt516*MW359211MW360924MW359470MW359722MW360210MW360437-MW360680Nebria cf. laevistriata Ledoux & RouxNIDMW359212MW360925MW359471MW359723MW360211MW360438MW359967MW360681Nebria livida (Linnaeus)717MW359213MW360926MW359472MW359724MW360212MW360439MW359968MW360682Nebria macrogona Bates717MW359214-MW359473MW359725MW360213MW360440MW359969MW360683Nebria martensi Huber & Schmidt16MW359215MW360927MW359474MW359726MW360214--MW360684Nebria numburica Huber & Schmidt16*MW359216MW360928MW359475MW359727MW360215MW360441MW359970MW360685Nebria oxyptera Daniel & Daniel16MW359217MW360929MW359476MW359728----Nebria pertinax Huber & Schmidt6JSMW359218MW360930MW359477MW359729MW360216MW360442MW359971MW360686Nebria pseudorestias Huber & Schmidt16MW359219MW360931MW359478MW359730MW360217MW360443MW359972MW360687Nebria sp YUN 16NIDMW359220MW360932MW359479MW359731MW360218MW360444MW359973MW360688Subgenus Nebria Psilonebria AndrewesNebria composita macra Ledoux & Roux917MW359221MW360933MW359480MW359732MW360219MW360445MW359974MW360689Nebria mentoincisa Huber & Schmidt821*MW359222MW360934MW359481MW359733MW360220MW360446MW359975MW360690Nebria nana Ledoux & Roux917MW359223MW360935MW359482MW359734MW360221MW360447MW359976MW360691Nebria pharina walteriana Ledoux & Roux817MW359224MW360936MW359483MW359735MW360222MW360448MW359977MW360692Nebria przewalskii Semenov917MW359225MW360937MW359484MW359736MW360223MW360449MW359978-Nebria roborowskii Semenov817MW359226MW360938MW359485MW359737MW360224MW360450MW359979MW360693Nebria cf. superna AndrewesJSMW359227MW360939MW359486MW359738MW360225MW360451MW359980MW360694Nebria sp XIZ 20NIDMW359228MW360940MW359487MW359739MW360226MW360452MW359981MW360695Subgenus Nebria Eurynebria GanglbauerNebria complanata (Linnaeus)8MW359229MW360941MW359488MW359740MW360227-MW359982MW360696Subgenus Nebria Eunebria JeannelNebria jockischii Sturm17MW359230MW360942MW359489MW359741MW360228MW360453MW359983MW360697Nebria lewisi Bates17MW359231MW360943MW359490MW359742MW360229MW360454MW359984MW360698Nebria limbigera Solsky17MW359232MW360944MW359491MW359743MW360230MW360455MW359985-Nebria morvani Ledoux & RouxJSMW359233MW360945MW359492MW359744MW360231MW360456MW359986MW360699Nebria perlonga Heyden17MW359234MW360946MW359493MW359745MW360232MW360457MW359987MW360700Nebria picicornis (Fabricius)17MW359235MW360947MW359494MW359746MW360233MW360458MW359988-Nebria psammodes (Rossi)17MW359236MW360948MW359495MW359747MW360234MW360459MW359989-Nebria uenoiana Habu17MW359237MW360949MW359496MW359748MW360235MW360460MW359990MW360701Nebria yunnana Bänninger17MW359238MW360950MW359497MW359749MW360236MW360461MW359991MW360702Nebria sp YUN 9CRCMW359239MW360951MW359498MW359750MW360237MW360462MW359992MW360703Nebria sp YUN 10-darkCRCMW359240MW360952MW359499MW359751MW360238MW360463MW359993MW360704Nebria sp YUN 10-paleCRCMW359241MW360953MW359500MW359752MW360239MW360464MW359994MW360705Nebria sp YUN 11CRCMW359242MW360954MW359501MW359753MW360240MW360465MW359995MW360706Catonebria SeriesReductonebria ComplexSubgenus Nebria Insulanebria New SubgenusNebria carbonaria Eschscholtz1018MW359243MW360955MW359502MW359754MW360241MW360466MW359996MW360707Nebria snowi Bates1018MW359244MW360956MW359503MW359755MW360242MW360467MW359997MW360708Subgenus Nebria Reductonebria ShilenkovNebria altaica Gebler18MW359245MW360957MW359504MW359756MW360243MW360468MW359998MW360709Nebria appalachia DarlingtonTMW359246MW360958MW359505MW359757MW360244MW360469MW359999MW360710Nebria chuskae KavanaughTMW359247MW360959MW359506MW359758MW360245MW360470MW360000MW360711Nebria darlingtoni KavanaughTMW359248MW360960MW359507MW359759MW360246MW360471MW360001MW360712Nebria desolata KavanaughTMW359249MW360961MW359508MW359760MW360247MW360472MW360002MW360713Nebria diversa LeConteTMW359250MW360962MW359509MW359761MW360248MW360473MW360003MW360714Nebria eschscholtzii Ménétriés1MW359251MW360963MW359510MW359762MW360249MW360474MW360004MW360715Nebria georgei KavanaughT*MW359252MW360964MW359511MW359763MW360250MW360475MW360005MW360716Nebria japonica Bates17MW359253MW360965MW359512MW359764MW360251MW360476MW360006MW360717Nebria mannerheimii Fischer von Waldheim1MW359254MW360966MW359513MW359765MW360252MW360477MW360007MW360718Nebria navajo KavanaughTMW359255MW360967MW359514MW359766MW360253MW360478MW360008MW360719Nebria obliqua LeConte COTMW359256MW360968MW359515MW359767MW360254MW360479MW360009MW360720Nebria obliqua LeConte UTTMW359257MW360969MW359516MW359768MW360255MW360480MW360010MW360721Nebria ochotica Sahlberg18MW359258MW360970MW359517MW359769MW360256MW360481MW360011MW360722Nebria pallipes Say1MW359259MW360971MW359518MW359770MW360257MW360482MW360012MW360723Nebria suturalis LeConte ABTMW359260MW360972MW359519MW359771MW360258MW360483MW360013MW360724Nebria suturalis LeConte COTMW359261MW360973MW359520MW359772MW360259MW360484MW360014MW360725Nebria suturalis LeConte NHTMW359262MW360974MW359521MW359773MW360260MW360485MW360015MW360726Subgenus Nebria Erwinebria New SubgenusNebria acuta Lindroth10TMW359263MW360975MW359522MW359774MW360261MW360486MW360016MW360727Nebria arkansana Casey10TMW359264MW360976MW359523MW359775MW360262MW360487MW360017MW360728Nebria charlottae Lindroth10TMW359265MW360977MW359524MW359776MW360263MW360488MW360018MW360729Nebria danmanni Kavanaugh10TMW359266MW360978MW359525MW359777MW360264MW360489MW360019MW360730Nebria edwardsi Kavanaugh10TMW359267MW360979MW359526MW359778MW360265MW360490MW360020MW360731Nebria fragilis Casey10TMW359268MW360980MW359527MW359779MW360266MW360491MW360021MW360732Nebria gregaria Fischer von Waldeim101MW359269MW360981MW359528MW359780MW360267MW360492MW360022MW360733Nebria haida Kavanaugh10TMW359270MW360982MW359529MW359781MW360268MW360493MW360023MW360734Nebria jeffreyi Kavanaugh10TMW359271MW360983MW359530MW359782MW360269MW360494MW360024MW360735Nebria lituyae Kavanaugh10TMW359272MW360984MW359531MW359783MW360270MW360495MW360025MW360736Nebria louiseae Kavanaugh10TMW359273MW360985MW359532MW359784MW360271MW360496MW360026MW360737Nebria lyelli Van Dyke10TMW359274MW360986MW359533MW359785MW360272MW360497MW360027MW360738Nebria modoc Kavanaugh10TMW359275MW360987MW359534MW359786MW360273MW360498MW360028MW360739Nebria oowah Kavanaugh10TMW359276MW360988MW359535MW359787MW360274MW360499MW360029MW360740Nebria quileute Kavanaugh10TMW359277MW360989MW359536MW359788MW360275MW360500MW360030MW360741Nebria sahlbergii Fischer von Waldheim10TMW359278MW360990MW359537MW359789MW360276MW360501MW360031MW360742Nebria sonorae Kavanaugh10TMW359279MW360991MW359538MW359790MW360277MW360502MW360032MW360743Nebria triad Kavanaugh10TMW359280MW360992MW359539MW359791MW360278MW360503MW360033MW360744Nebria wallowae Kavanaugh10TMW359281MW360993MW359540MW359792MW360279MW360504MW360034MW360745Nebria zioni Van Dyke10TMW359282MW360994MW359541MW359793MW360280MW360505MW360035MW360746Catonebria ComplexSubgenus Nebria Nivalonebria New SubgenusNebria paradisi Darlington OR11TMW359283MW360995MW359542MW359794MW360281MW360506MW360036MW360747Nebria paradisi Darlington WA11TMW359284MW360996MW359543MW359795MW360282MW360507MW360037MW360748Nebria turmaduodecima Kavanaugh11TMW359285MW360997MW359544MW359796MW360283MW360508MW360038MW360749Subgenus Nebria Apteronebria New SubgenusNebria balli Kavanaugh12TMW359286MW360998MW359545MW359797MW360284MW360509MW360039MW360750Nebria carri Kavanaugh12TMW359287MW360999MW359546MW359798MW360285MW360510MW360040MW360751Nebria kincaidi Schwarz12TMW359288MW361000MW359547MW359799MW360286MW360511MW360041MW360752Nebria ovipennis LeConte12TMW359289MW361001MW359548MW359800MW360287MW360512MW360042MW360753Nebria sierrae Kavanaugh12TMW359290MW361002MW359549MW359801MW360288MW360513MW360043MW360754Nebria spatulata Van Dyke12TMW359291MW361003MW359550MW359802MW360289MW360514MW360044MW360755Subgenus Nebria Palapteronebria New SubgenusNebria arinae Dudko & Shilenkov12RDMW359292MW361004MW359551MW359803MW360290MW360515MW360045MW360756Nebria baenningeri baenningeri Dudko & Shilenkov12RDMW359293MW361005MW359552MW359804MW360291MW360516MW360046MW360757Nebria baenningeri korgonica Dudko & Shilenkov12RDMW359294MW361006MW359553MW359805MW360292MW360517MW360047MW360758Nebria lyubechanskii Dudko12RDMW359295MW361007MW359554MW359806MW360293MW360518MW360048MW360759Nebria mellyi mellyi Gebler12RDMW359296MW361008MW359555MW359807MW360294MW360519MW360049MW360760Nebria mellyi teletskiana Dudko & Shilenkov12RDMW359298MW361010MW359557MW359809MW360296MW360521MW360051MW360762Nebria roddi Dudko & Shilenkov12RDMW359297MW361009MW359556MW359808MW360295MW360520MW360050MW360761Nebria sajana dubatolovi Dudko & Shilenkov12RDMW359299MW361011MW359558MW359810MW360297MW360522MW360052MW360763Nebria sajana sajana Dudko & Shilenkov12RDMW359300MW361012MW359559MW359811MW360298MW360523MW360053MW360764Nebria sajana sarlyk Dudko & Shilenkov12RDMW359301MW361013MW359560MW359812MW360299MW360524MW360054MW360765Nebria sajana sitnikovi Dudko & Shilenkov12RDMW359302MW361014MW359561MW359813MW360300MW360525MW360055MW360766Subgenus Nebria Catonebria ShilenkovNebria aenea GeblerRDMW359303MW361015MW359562MW359814MW360301MW360526MW360056MW360767Nebria albimontis KavanaughTMW359304MW361016MW359563MW359815MW360302MW360527MW360057MW360768Nebria banksii Crotch22MW359305MW361017MW359564MW359816MW360303MW360528MW360058MW360769Nebria baumanni KavanaughTMW359306MW361018MW359565MW359817MW360304MW360529MW360059MW360770Nebria beverlianna KavanaughTMW359307MW361019MW359566MW359818MW360305MW360530MW360060MW360771Nebria calva KavanaughTMW359308MW361020MW359567MW359819MW360306MW360531MW360061MW360772Nebria cascadensi s KavanaughTMW359309MW361021MW359568MW359820MW360307MW360532MW360062MW360773Nebria catenata CaseyTMW359310MW360812MW359569MW359821MW360308MW360533MW360063MW360774Nebria catenulata Fischer von Waldheim22MW359311MW361022MW359570MW359822MW360309MW360534MW360064MW360775Nebria coloradensis Van DykeTMW359312MW361023MW359571MW359823MW360310MW360535MW360065MW360776Nebria fragariae KavanaughTMW359313MW361024MW359572MW359824MW360311MW360536MW360066MW360777Nebria fulgida Gebler22MW359314MW361025MW359573MW359825MW360312MW360537MW360067MW360778Nebria gebleri DejeanTMW359315MW361026MW359574MW359826MW360313MW360538MW360068MW360779Nebria giulianii KavanaughTMW359316MW361027MW359575MW359827MW360314MW360539MW360069MW360780Nebria holzunensis Dudko & Shilenkov22MW359317MW361028MW359576MW359828MW360315MW360540MW360070MW360781Nebria ingens HornTMW359318MW361029MW359577MW359829MW360316MW360541MW360071MW360782Nebria labontei KavanaughTMW359319MW361030MW359578MW359830MW360317MW360542MW360072MW360783Nebria lamarckensis KavanaughTMW359320MW361031MW359579MW359831MW360318MW360543MW360073MW360784Nebria meanyi Van Dyke CATMW359321MW361032MW359580MW359832MW360319MW360544MW360074MW360785Nebria meanyi Van Dyke WATMW359322MW361033MW359581MW359833MW360320MW360545MW360075MW360786Nebria metallica Fischer von Waldheim1MW359323MW361034MW359582MW359834MW360321MW360546MW360076MW360787Nebria pasquineli KavanaughTMW359324MW361035MW359583MW359835MW360322MW360547MW360077MW360788Nebria pektusanica Horvatovich22MW359325MW361036MW359584MW359836MW360323MW360548MW360078MW360789Nebria piperi Van DykeTMW359326MW361037MW359585MW359837MW360324MW360549MW360079MW360790Nebria piute Erwin & BallTMW359327MW360813MW359586MW359838MW360325MW360550MW360080MW360791Nebria praedicta Kavanaugh & SchovilleTMW359328MW361038MW359587MW359839MW360326MW360551MW360081MW360792Nebria purpurata LeConte COTMW359329MW361039MW359588MW359840MW360327MW360552MW360082MW360793Nebria purpurata LeConte NMTMW359330MW361040MW359589MW359841MW360328MW360553MW360083MW360794Nebria rathvoni LeConteTMW359331MW361041MW359590MW359842MW360329MW360554MW360084MW360795Nebria riversi Van DykeTMW359332MW361042MW359591MW359843MW360330MW360555MW360085MW360796Nebria schwarzi Van DykeTMW359333MW361043MW359592MW359844MW360331MW360556MW360086MW360797Nebria sevieri KavanaughTMW359334MW361044MW359593MW359845MW360332MW360557MW360087MW360798Nebria sierrablancae KavanaughTMW359335MW361045MW359594MW359846MW360333MW360558MW360088MW360799Nebria siskiyouensis KavanaughTMW359336MW361046MW359595MW359847MW360334MW360559MW360089MW360800Nebria splendida Fischer von Waldheim22MW359337MW361047MW359596MW359848MW360335MW360560MW360090MW360801Nebria steensensis KavanaughTMW359338MW361048MW359597MW359849MW360336MW360561MW360091MW360802Nebria sylvatica KavanaughTMW359339MW361049MW359598MW359850MW360337MW360562MW360092MW360803Nebria trifaria LeConteTMW359340MW361050MW359599MW359851MW360338MW360563MW360093MW360804Nebria utahensis KavanaughTMW359341MW361051MW359600MW359852MW360339MW360564MW360094MW360805Nebria vandykei Bänninger1MW359342MW361052MW359601MW359853MW360340MW360565MW360095MW360806Nebria wyeast KavanaughTMW359343MW361053MW359602MW359854MW360341MW360566MW360096MW360807Outgroup taxaTrachypachidaeTrachypachus inermis Motschulsky1MW359344MW361054MW359603MW359855MW360342--MW360808Trachypachus slevini Van DykeTMW359345MW361055MW359604MW359856DHK0513a--MW360809CarabidaeCarabiniCalosoma scrutator (Fabricius)1EU677684MW361056MW359605MW359857EU677530MW360567EU677634EU677661CychriniScaphinotus petersi RoeschkeDREU658920MW361058MW359607MW359859EU677531MW360569EU677635EU658922MetriiniMetrius contractus Eschscholtz1MW359346MW361059MW359608MW359860MW360343MW360570MW360097MW360810LoriceriniLoricera pilicornis (Fabricius)1MW359347MW361060MW359609MW359861MW360344MW360571MW360098MW360811BembidiiniAsaphidion yukonense WickhamDMEU677686---EU677540-EU677638EU677666Bembidion antiquum DejeanDMEF648850MW361061EF649127.1EF649127.1EF649402MW360572MW360099EF649489PterostichiniPterostichus melanarius (Illiger)DMAF398707MW361062MW359610MW359862EU677533MW360573MW360100AF398623
1 Formerly assigned to subgenus Pterostichus Epinebriola. 2 Formerly assigned to subgenus Pterostichus Marggia. 3 Formerly assigned to subgenus Pterostichus Nebriorites. 4 Formerly assigned to subgenus Pterostichus Pseudonebriola. 5 Formerly assigned to subgenus Pterostichus Barbonebriola. 6 Formerly assigned to subgenus Pterostichus Patrobonebria. 7 Formerly assigned to subgenus Pterostichus Paranebria. 8 Formerly assigned to subgenus Pterostichus Asionebria. 9 Formerly assigned to subgenus Pterostichus Eunebria. 10 Formerly assigned to subgenus Pterostichus Reductonebria. 11 Formerly assigned to subgenus Pterostichus Nakanebria. 12 Formerly assigned to subgenus Pterostichus Catonebria.