Temporal evolution of the asymmetry of links, cophenetic correlation coefficients (CCC), maximum modularities (Max. Mod.), and consecutive cluster determinations for the metapopulation.

Statistically significant results are presented in bold. For the asymmetry tests, “»” indicate the direction of the asymmetry. Codes for the populations are: W for West, S for South, C for Centre, NE for North-East and NW for North-West.