TABLE 3 in Amphisbaena lumbricalis Vanzolini, 1996 is a synonym of Amphisbaena carvalhoi Gans, 1965 (Squamata, Amphisbaenidae)

TABLE 3. Morphological comparisons among four-pored Amphisbaena species from South America. Pores: N = number of precloacal pores; MH = median hiatus (Y = present, N = absent). Head: SL = number of supralabials; IL = number of infralabials; PG = number of postgenial rows; M = pair of enlarged malar scales (Y = present, N = absent); PM = postmalars (Y = present, N = absent). Body and tail: BA = body annuli; LA = lateral annuli; CA = caudal annuli; DS = dorsal segments at midbody annulus; VS = ventral segments at midbody annulus; PeC = precloacal segments; PoC = postcloacal segments; AA = autotomic annulus (N = autotomic site absent); TT = tail tip (R = rounded; LC = laterally compressed; VK = vertically keeled). Max SVL = maximum snout-vent length (in millimeters). Blank cells indicate information not provided by the consulted literature.
SpeciesPores N MHSLILHead PGMPMBALACABody and tail DS VSPeCPoCAATTMax SVLSourceA. carvalhoi4 (rarely 2, 5, 6)N331–2YN223–2493–419–2612–1416–206–910–146–10R168.5This studyA. acangaoba4–8N431–2YY216–2392–413–1718–2418–246–1013–18N29044A. alba4–10N431–2YY198–24813–2130–4235–46NR5502A. albocingulata4N331–2YN190–2043–424–2712–1416–186–810–127–913830,43A. angustifrons3–6N3–432YY190–2182–412–1820–3120–306–1012–20NLC34510,21A. arda4431YY2423023238828527A. arenaria4Y332YN285–30722–2312–1414–16810–126–7R24518,38A. arenicola4NH3–432HYHY199–21620–2212–1416–186H11H8–9R16843A. bahiana4N332YY204–22314–1712–1614–16NVK21541A. bedai4Y431YHN272–28422–2318–2016–18630016A. bolivica3–6N432YY200–23118–2627–3626–363–5R32810,20,24A. borelli4Y332N239–26117–1914–1816–206–8VK8,33A. brasiliana4Y3–431YY213–2292–411–1518–2118–218–1010–17NR31712A. caetitensis4N332YN186–19410–121619–226–712–14NPT23540A. camura3–6N432YY188–20714–1928–4228–453–5R38010,20,24A. cegei0 or 4N332YN179–1993–421–2417–2220–24613–156–8R11125,33A. cuiabana4Y3332YN278–3093–517–201414–166–811–138–10VK31626,29A. cunhai4N331YY224–2393–425–2614–1614–1867–115–7R24623A. darwinii2–5N332YY174–1992–418–2513–1916–236–89–156–10LC30411,43A. elbakyanae4N331NY245–257420–2413–1516–186–711–126–8R23745A. frontalis0, 4 or 6N332–3YN235–2752–423–2914–1814–165–814–185–728346A. gracilis4N332NY224–248521–2413–1614–17612–136–7R25214,45A. hastata4N332–3YN266–27344035–371812–16R14917A. heathi0–4N332YY182–2013–423–2811–1218–20811–127–9R1369,37A. heterozonata2–6N331–2YY189–20718–2313–1816–2268–165–8LC27711,33,43A. hogei4N33Y177–19115–1910–1314–1812–1613443A. hoogmoedi4N332YN247–2522722–2420H10H12H7–8R29142A. medemi4N332YN230–2353–517–1814–1617–186–812–175–7R218422A. metallurga2–4N331–2YY–N185–1992–323–2512–1414–166–79–167–9R17932,35A. mongoyo4N332YN208251416610R14139A. munoai4N331–2YY202–21818–2310–1413–186–87–145–9R15143A. myersi433221281616820A. nana4NH332HYHY195–21618–2212–1414–176H13H7–10R15243A. nigricauda0, 4 or 5N332YN192–2263–519–249–1113–16610–136–10R16811,31A. occidentalis4N432YY262–2753–518–2116–2024–266–911–16?R2431A. pericensis4N332YN198–2183–516–1912–1616–206–811–146–8LC1535A. plumbea4N431–2YY210–2823–616–2118–2720–306–912–165–9R36521A. polygrammica4331Y270221826?33A. prunicolor4N331–2YY181–21518–2710–1714–206–87–147–11R23830,43A. ridleyi4N431–2YY172–1922–414–1716–1820–286–811–14NR2456A. rozei4N331YY205–20932015–16146H14H625234A. sanctaeritae4N331–2YY269–2882–318–20101466–7LC16519,35A. slateri4N331–2YY–N176–2133–420–2410–1414–166–810–127–10R16333A. slevini4N22NNN204–2112–423–2510–1410–126–89–124–6R1154A. spurrelli4N331–2YY218–2223–518–2016–1816–1872973A. steindachneri4Y331–2N256–2662–517–1812–1612–166–1010–197VK2568,33A. supernumeraria4N332YN333–33722–231417–1810–12VK19528A. talisiae4N331YN205–2342–417–2910–1414–185–88–146–8R14636A. tiaraju4NH331HYHY204–22118–2510–1413–166H10H7–9R18343A. townsendi4N431YY261–2792–522–2616–1922–286–911–167–8R2301A. trachura3–4N331–2YY186–20815–2514–2116–246–89–145–932211,43A. tragorrhectes4N44341YY196311212611015A. vanzolinii4N221NN200–2314–528–3112–1612–186127–141537,45A. vermicularis4 (rarely 5 or 6)N431–2YY211–2543–523–3018–2618–256–910–165–7R32713
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1 Total number of segments around midbody (absence of lateral sulcus). 2 Tail rounded, with slight distal pinching. 3 Amphisbaena cuiabana has a median hiatus in the series of precloacal pores that results from the position of median pores near the lateral sutures of wide wedge-shaped segments; non-pored medial segments are absent. 4 Possibly three supralabials and two infralabials, not four supralabials and three infralabials as stated by Vanzolini (1971). H Information based only on the holotype, not the entire type series. Source: 1 = Gans (1961), 2 = Gans (1962a), 3 = Gans (1962b), 4 = Gans (1963a), 5 = Gans (1963b), 6 = Gans (1963c), 7 = Gans (1963d), 8 = Gans (1964), 9 = Gans (1965a), 10 = Gans (1965b), 11 = Gans (1966), 12 = Gans (1971), 13 = Gans & Amdur 1966, 14 = Gonzalez-Sponga & Gans (1971), 15 = Vanzolini (1971), 16 = Vanzolini (1991a), 17 = Vanzolini (1991b), 18 = Vanzolini (1991c), 19 = Vanzolini (1994), 20 = Vanzolini (2002), 21 = Gans & Diefenbach (1972), 22 = Gans & Mathers (1977), 23 = Hoogmoed & Ávila-Pires (1991), 24 = Montero (1996), 25 = Montero et al. (1997), 26 = Strüssmann & Carvalho (2001), 27 = Rodrigues (2003), 28 = Mott et al. (2009), 29 = Silva et al. (2010), 30 = Perez et al. (2012), 31 = Sousa e Lima et al. (2014), 32 = Costa et al. (2015), 33 = Costa et al. (2018a), 34 = Costa et al. (2018b), 35 = Costa et al. (2019a), 36 = Costa et al. (2019b), 37 = Lisboa et al. (2016), 38 = Teixeira Jr. et al. (2016), 39 = Teixeira Jr. et al. (2019), 40 = Almeida et al. (2018), 41 = Dal Vechio et al. (2018), 42 = Oliveira et al. (2018), 43 = Perez & Borges-Martins (2019), 44 = Ribeiro et al. (2020), 45 = Torrez-Ramírez et al. (2021), 46 = Ribeiro-Junior et al. (2022).