DyHDER: Dynamic Habitat Disturbance and Ecological Resilience (“Die Harder”) - a metapopulation matrix model incorporating subpopulation habitat conditions and physical connectivity into a population viability analysis (PVA) framework.

DyHDER (pronounced “Die Harder”) is a spatially explicit, stage-structured matrix population model. The model can be used in a PVA framework, can be applied to metapopulations of any size or species, and was specifically designed with a structure capable of evaluating mechanistic metapopulation responses to habitat condition (e.g., disturbance). This is accomplished through the use of species-specific habitat suitability relations. DyHDER also includes a metapopulation dispersal model that drives probabilistic emigration and immigration rates between subpopulation sites as a function of changing habitat conditions and site densities. DyHDER is intended to provide a framework for evaluating ecological population-level responses of probabilistic or theoretical habitat disturbance scenarios (e.g., flood, drought, wildfire, habitat restoration, fragmentation) of varying location, magnitude, and duration.