Free parameters of the space-based and mixed models and their values which were determined by evolutionary optimization.

Columns “Fit (last gen.)” show the mean and standard deviation across all 32 parameter sets from the last generation of the evolutionary optimization (for a plot of the full parameter space, see the S3–S7 Figs). Columns “Fit (best)” show the parameter values that lead to the best fitness among all individuals and which are the parameter values used for the model comparison shown in Fig 4. The parameter for the size of the oculomotor drift is set to σD = 0.125 dva for all models.