Features of the right tarsometatarsus of UNSM 20030 that were identified by a phylogenetic analysis (see [8]) as uniting Fumicollis hoffmani, Parahesperornis, and Hesperornis in a clade.

A, well-developed depression (d) on the lateral face of the proximal end of metatarsal IV; B, prominent grooves (g) separating the metatarsals along the entire length of shaft and ridge (r) that forms the dorsal surface of metatarsal IV; C, the enclosed distal vascular foramen (df) between the trochlea of metatarsals III and IV is tear-drop shaped in plantar view (note the trochleae of the specimen have been labelled incorrectly); D, metatarsals IV, III, and II progressively displaced plantarly (shingled; displacement shown with arrows); and E, shaft of metatarsal IV is widest at midshaft and tapers at both the proximal and distal ends (widths indicated with arrows). Abbreviations: d, depression; df, distal foramen; g, groove; r, ridge; II–IV, metatarsal II–IV.