Pearson's correlation coefficients for comparisons between biological and physical indices and smolt-to-adult return ratios (SAR).

FL = fork length (mm). M = mass (g). MEBT is the annual mean day of marine entry in relation to the biological spring transition whereas MEPT is the annual mean day of marine entry in relation to the physical spring transition (see text for additional details). Subscripts indicate the months over which data were averaged (e.g., PDO7_9 = mean PDO from July to September). Adjusted significant values (†P